Species in mammal order Afrosoricida
Afrosoricida is an order of placental mammals . Members of this order are called afrosoricids, and include golden moles , otter shrews , and tenrecs . They are found in Africa, generally in forests, but also inland wetlands , shrublands , and grasslands . They range in size from the least shrew tenrec , at 4 cm (2 in) plus a 6 cm (2 in) tail, to the giant otter shrew , at 34 cm (13 in) plus a 39 cm (15 in) tail. Afrosoricids primarily eat invertebrates , particularly insects and earthworms, though some will also eat small lizards or other vertebrates. The golden moles have vestigial eyes covered with skin, and track their prey through vibrations rather than sight.[ 1] No population estimates have been made for any afrosoricid species, though the De Winton's golden mole is classified as Critically Endangered and the giant golden mole , Gunning's golden mole , Jenkins's shrew tenrec , Juliana's golden mole , Marley's golden mole , northern shrew tenrec , and Van Zyl's golden mole are categorized as endangered species .
The fifty-five extant species of Afrosoricida are divided into two suborders , with Chrysochloridea containing the family Chrysochloridae , or golden moles, and Tenrecomorpha containing the families Potamogalidae , or otter shrews, and Tenrecidae , or tenrecs. Chrysochloridae is split into the subfamilies Chrysochlorinae, containing eleven species in six genera , and Amblysominae, containing ten species in four genera. Potamogalidae consists of three species in two genera, while Tenrecidae contains the subfamilies of Geogalinae , comprising a single species, Oryzorictinae , containing twenty-five species in three genera, and Tenrecinae , containing five species in four genera. The order as a whole was traditionally grouped with the hedgehogs, shrews, and moles as part of the order Lipotyphla , but modern molecular phylogenetic analysis resulted in that order being split into Afrosoricida and Eulipotyphla . Few extinct Afrosoricida species have been discovered, though due to ongoing research and discoveries the exact number and categorization are not fixed.[ 2]
Conventions
Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species . Range maps are provided wherever possible; if a range map is not available, a description of the afrosoricid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. All extinct species or subspecies listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "†".
Classification
The order Afrosoricida consists of two suborders , Chrysochloridea and Tenrecomorpha . Chrysochloridea consists of the family Chrysochloridae , or golden moles, and Tenrecomorpha contains the families Potamogalidae , or otter shrews, and Tenrecidae , or tenrecs. Chrysochloridae contains twenty-one species in ten genera , divided into two subfamilies. Potamogalidae consists of three species in two genera, while Tenrecidae contains thirty-one species in eight genera, divided into three subfamilies. Many of these species are further subdivided into subspecies. This does not include hybrid species or extinct prehistoric species.
Afrosoricids
The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis , as supported by both the IUCN and the American Society of Mammalogists .[ 6]
Suborder Chrysochloridea
Family Chrysochloridae
Subfamily Chrysochlorinae
Genus Chrysochloris – Lacépède , 1799 – three species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Cape golden mole
C. asiatica (Linnaeus , 1758)
Western South Africa
Size : 9–12 cm (4–5 in) long[ 13] Habitat : Forest, shrubland, and grassland[ 14] Diet : Insects and small invertebrates , as well as small lizards[ 15]
LC
Unknown [ 14]
Stuhlmann's golden mole
C. stuhlmanni Matschie , 1894
C. s. balsaci
C. s. stuhlmanni
C. s. stuhlmanni
Scattered central Africa
Size : 10–13 cm (4–5 in) long[ 16] Habitat : Forest, shrubland, and grassland[ 17] Diet : Earthworms and insect larvae, as well as other invertebrates[ 16]
LC
Unknown [ 17]
Visagie's golden mole
C. visagiei Broom , 1950
Western South Africa
Size : About 10 cm (4 in) long[ 18] Habitat : Shrubland[ 19] Diet : Unknown[ 18]
DD
Unknown [ 19]
Genus Chrysospalax – Gill , 1883 – two species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Giant golden mole
C. trevelyani (Günther , 1875)
Southern South Africa
Size : 20–24 cm (8–9 in) long[ 20] Habitat : Forest and grassland[ 21] Diet : Small invertebrates, especially giant earthworms and millipedes, as well as small vertebrates[ 21]
EN
Unknown [ 21]
Rough-haired golden mole
C. villosus (Smith , 1833)
C. v. dobsoni
C. v. leschae
C. v. rufopallidus
C. v. rufus
C. v. transvaalensis
C. v. villosus
South Africa
Size : 12–18 cm (5–7 in) long[ 22] Habitat : Shrubland and grassland[ 23] Diet : Small invertebrates, especially termites and earthworms[ 22]
VU
Unknown [ 23]
Genus Eremitalpa – Roberts, 1924 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Grant's golden mole
E. granti (Broom , 1907)
E. g. granti
E. g. namibensis
Western South Africa and Namibia
Size : 7–9 cm (3–4 in) long[ 28] Habitat : Shrubland and desert[ 29] Diet : Termites and other insects, as well as roots and small lizards[ 28]
LC
Unknown [ 29]
Subfamily Amblysominae
Genus Amblysomus – Pomel , 1848 – five species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Fynbos golden mole
A. corriae Thomas , 1905
A. c. corriae
A. c. devilliersi
Southern South Africa
Size : 10–13 cm (4–5 in) long[ 30] Habitat : Forest, savanna, shrubland, and grassland[ 31] Diet : Insects[ 30]
NT
Unknown [ 31]
Highveld golden mole
A. septentrionalis Roberts , 1913
Eastern South Africa
Size : 10–15 cm (4–6 in) long[ 32] Habitat : Forest, savanna, shrubland, and grassland[ 33] Diet : Earthworms and other invertebrates[ 32]
NT
Unknown [ 33]
Hottentot golden mole
A. hottentotus (Smith , 1829)
A. h. hottentotus
A. h. iris (Zulu golden mole )
A. h. longiceps
A. h. meesteri
A. h. pondoliae
South Africa
Size : 10–14 cm (4–6 in) long[ 34] Habitat : Forest, savanna, shrubland, and grassland[ 35] Diet : Earthworms and other invertebrates[ 34]
LC
Unknown [ 35]
Marley's golden mole
A. marleyi Roberts , 1931
Eastern South Africa
Size : 9–12 cm (4–5 in) long[ 36] Habitat : Forest, savanna, shrubland, and grassland[ 37] Diet : Insects[ 36]
EN
Unknown [ 37]
Robust golden mole
A. robustus Bronner , 2000
Eastern South Africa
Size : 10–15 cm (4–6 in) long[ 38] Habitat : Shrubland and grassland[ 39] Diet : Small insects and earthworms[ 38]
VU
Unknown [ 39]
Genus Calcochloris – Mivart , 1867 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Yellow golden mole
C. obtusirostris (Peters , 1851)
C. o. chrysillus
C. o. limpopoensis
C. o. obtusirostris
Mozambique, South Africa, and Zimbabwe
Size : 8–11 cm (3–4 in) long[ 40] Habitat : Forest and savanna[ 41] Diet : Insects as well as small lizards[ 40]
LC
Unknown [ 41]
Suborder Tenrecomorpha
Family Potamogalidae
Genus Micropotamogale – Heim de Balsac , 1954 – two species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Nimba otter shrew
M. lamottei Heim de Balsac , 1954
Mount Richard-Molard area in western Africa
Size : 12–16 cm (5–6 in) long, plus 9–14 cm (4–6 in) tail[ 50] Habitat : Forest and inland wetlands[ 51] Diet : Crabs and catfish, as well as insects and tadpoles[ 51]
VU
Unknown [ 51]
Ruwenzori otter shrew
M. ruwenzorii (de Witte , Frechkop , 1955)
Central Africa
Size : 12–20 cm (5–8 in) long, plus 10–15 cm (4–6 in) tail[ 52] Habitat : Forest and inland wetlands[ 53] Diet : Insect larvae and worms, as well as small fish, frogs, and crabs[ 52]
LC
Unknown [ 53]
Genus Potamogale – Du Chaillu , 1860 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Giant otter shrew
P. velox (Du Chaillu , 1860)
Central Africa
Size : 30–34 cm (12–13 in) long, plus 23–29 cm (9–11 in) tail[ 54] Habitat : Forest and inland wetlands[ 55] Diet : Fishes, crabs, shrimps, and water insects, as well as frogs[ 55]
LC
Unknown [ 55]
Family Tenrecidae
Subfamily Geogalinae
Subfamily Oryzorictinae
Genus Microgale – Thomas , 1882 – twenty-one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Cowan's shrew tenrec
M. cowani Thomas , 1882
Madagascar
Size : 6–10 cm (2–4 in) long, plus 6–8 cm (2–3 in) tail[ 58] Habitat : Forest[ 59] Diet : Insects and other invertebrates, as well as small vertebrates[ 60]
LC
Unknown [ 59]
Drouhard's shrew tenrec
M. drouhardi Grandidier , 1934
Eastern Madagascar
Size : 6–9 cm (2–4 in) long, plus 5–9 cm (2–4 in) tail[ 61] Habitat : Forest[ 62] Diet : Insects and other invertebrates, as well as small vertebrates[ 60]
LC
Unknown [ 62]
Dryad shrew tenrec
M. dryas Jenkins , 1992
Northeastern Madagascar
Size : 17–18 cm (7–7 in) long, plus tail[ 63] Habitat : Forest[ 64] Diet : Invertebrates[ 63]
VU
Unknown [ 64]
Gracile shrew tenrec
M. gracilis (Major , 1896)
Eastern Madagascar
Size : 16–19 cm (6–7 in) long, plus tail[ 65] Habitat : Forest[ 66] Diet : Believed to be insects, earthworms, and other invertebrates[ 65]
LC
Unknown [ 66]
Grandidier's shrew tenrec
M. grandidieri Olson , Rakotomalala , Hildebrandt , Lanier , Raxworthy , Goodman , 2009
Western Madagascar
Size : 5–8 cm (2–3 in) long, plus 3–4 cm (1–2 in) tail[ 67] Habitat : Forest and shrubland[ 68] Diet : Believed to be insects and other invertebrates[ 67]
LC
Unknown [ 68]
Greater long-tailed shrew tenrec
M. principula Thomas , 1926
Eastern Madagascar
Size : 7–8 cm (3 in) long, plus 14–18 cm (6–7 in) tail[ 69] Habitat : Forest[ 70] Diet : Insects and other invertebrates, as well as small vertebrates[ 60]
LC
Unknown [ 70]
Jenkins's shrew tenrec
M. jenkinsae Goodman , Soarimalala , 2004
Southwestern Madagascar
Size : 7–8 cm (3 in) long, plus 7–9 cm (3–4 in) tail[ 71] Habitat : Forest and shrubland[ 72] Diet : Insects and other invertebrates[ 60]
EN
Unknown [ 72]
Least shrew tenrec
M. pusilla Major , 1896
Eastern Madagascar
Size : 4–6 cm (2 in) long, plus 6–8 cm (2–3 in) tail[ 73] Habitat : Forest and inland wetlands[ 74] Diet : Insects and other invertebrates[ 60]
LC
Unknown [ 74]
Lesser long-tailed shrew tenrec
M. longicaudata Thomas , 1882
Central and northern Madagascar
Size : 6–8 cm (2–3 in) long, plus 11–16 cm (4–6 in) tail[ 60] Habitat : Forest[ 75] Diet : Insects, arachnids, crustaceans, and other invertebrates[ 76]
LC
Unknown [ 75]
Major's long-tailed tenrec
M. majori Thomas , 1918
Madagascar
Size : 5–7 cm (2–3 in) long, plus 10–14 cm (4–6 in) tail[ 77] Habitat : Forest[ 78] Diet : Insects and other invertebrates[ 60]
LC
Unknown [ 78]
Montane shrew tenrec
M. monticola Goodman , Jenkins , 1998
Northeastern Madagascar
Size : 7–10 cm (3–4 in) long, plus 9–12 cm (4–5 in) tail[ 79] Habitat : Forest[ 80] Diet : Insects and other invertebrates[ 60]
VU
Unknown [ 80]
Naked-nosed shrew tenrec
M. gymnorhyncha Jenkins , Goodman , Raxworthy , 1996
Eastern Madagascar
Size : 13–18 cm (5–7 in) long, plus tail[ 81] Habitat : Forest[ 82] Diet : Insects, as well as small mammals, amphibians, vegetation, and potentially carrion[ 81]
LC
Unknown [ 82]
Nasolo's shrew tenrec
M. nasoloi Jenkins , 1999
Western Madagascar
Size : About 8 cm (3 in) long, plus 5 cm (2 in) tail[ 83] Habitat : Forest[ 84] Diet : Insects and other invertebrates[ 60]
VU
Unknown [ 84]
Northern shrew tenrec
M. jobihely Goodman , Raxworthy , Maminirina , Olson , 2006
Eastern and northern Madagascar
Size : 4–7 cm (2–3 in) long, plus 4–6 cm (2 in) tail[ 85] Habitat : Forest[ 86] Diet : Insects and other invertebrates[ 60]
EN
Unknown [ 86]
Pale shrew tenrec
M. fotsifotsy Jenkins , Raxworthy , Nussbaum , 1997
Eastern and northern Madagascar
Size : 6–9 cm (2–4 in) long, plus 7–10 cm (3–4 in) tail[ 79] Habitat : Forest[ 87] Diet : Insects and other invertebrates[ 60]
LC
Unknown [ 87]
Pygmy shrew tenrec
M. parvula Grandidier , 1934
Eastern and northern Madagascar
Size : 5–7 cm (2–3 in) long, plus 4–7 cm (2–3 in) tail[ 88] Habitat : Forest[ 89] Diet : Insects and other invertebrates[ 60]
LC
Unknown [ 89]
Short-tailed shrew tenrec
M. brevicaudata Grandidier , 1899
Western and northern Madagascar
Size : 6–7 cm (2–3 in) long, plus 3 cm (1 in) tail[ 90] Habitat : Forest and shrubland[ 91] Diet : Insects and small vertebrates[ 90]
LC
Unknown [ 91]
Shrew-toothed shrew tenrec
M. soricoides Jenkins , 1993
Eastern Madagascar
Size : 7–11 cm (3–4 in) long, plus 8–11 cm (3–4 in) tail[ 92] Habitat : Forest[ 93] Diet : Insects and other invertebrates, as well as small vertebrates[ 60]
LC
Unknown [ 93]
Taiva shrew tenrec
M. taiva Major , 1896
Eastern Madagascar
Size : 5–8 cm (2–3 in) long, plus 7–9 cm (3–4 in) tail[ 85] Habitat : Forest[ 94] Diet : Insects and other invertebrates[ 60]
LC
Unknown [ 94]
Thomas's shrew tenrec
M. thomasi Major , 1896
Eastern Madagascar
Size : 7–12 cm (3–5 in) long, plus 5–8 cm (2–3 in) tail[ 77] Habitat : Forest[ 95] Diet : Insects and other invertebrates[ 60]
LC
Unknown [ 95]
Web-footed tenrec
M. mergulus Major , 1896
Eastern Madagascar
Size : 12–17 cm (5–7 in) long, plus 11–17 cm (4–7 in) tailHabitat : Forest and inland wetlandsDiet : Insects, as well as tadpoles and crayfish
VU
Unknown
Genus Nesogale – Thomas , 1918 – two species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Dobson's shrew tenrec
N. dobsoni (Thomas , 1884)
Eastern and northern Madagascar
Size : 9–12 cm (4–5 in) long, plus 10–11 cm (4 in) tail[ 98] Habitat : Forest[ 99] Diet : Insects and ant eggs[ 98]
LC
Unknown [ 99]
Talazac's shrew tenrec
N. talazaci (Major , 1896)
Eastern Madagascar
Size : 4–13 cm (2–5 in) long, plus 4–16 cm (2–6 in) tail[ 100] Habitat : Forest[ 101] Diet : Insects as well as frogs[ 100]
LC
Unknown [ 101]
Genus Oryzorictes – Grandidier , 1870 – two species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Four-toed rice tenrec
O. tetradactylus Milne-Edwards , Grandidier , 1882
Southeastern Madagascar
Size : 10–12 cm (4–5 in) long, plus 4–6 cm (2 in) tail[ 102] Habitat : Forest, shrubland, grassland, and inland wetlands[ 103] Diet : Worms, insects, and other invertebrates[ 102]
DD
Unknown [ 103]
Mole-like rice tenrec
O. hova Grandidier , 1870
Eastern and northern Madagascar
Size : 9–13 cm (4–5 in) long, plus tail[ 104] Habitat : Forest and inland wetlands[ 105] Diet : insects and earthworms, as well as plants[ 104]
LC
Unknown [ 105]
Subfamily Tenrecinae
Genus Echinops – Linnaeus , 1753 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Lesser hedgehog tenrec
E. telfairi Martin , 1838
Southwestern Madagascar
Size : 14–18 cm (6–7 in) long, plus 13–17 cm (5–7 in) tail[ 106] Habitat : Forest, savanna, shrubland, and grassland[ 107] Diet : Insects and fruit[ 106]
LC
Unknown [ 107]
Genus Setifer – (Froriep , 1806) – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Greater hedgehog tenrec
S. setosus (Schreber , 1778)
Madagascar
Size : 16–23 cm (6–9 in) long, plus tail[ 113] Habitat : Forest, savanna, shrubland, and grassland[ 114] Diet : Insects, grubs, other invertebrates, and fruit[ 113]
LC
Unknown [ 114]
Genus Tenrec – Lacépède , 1799 – one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Tailless tenrec
T. ecaudatus (Schreber , 1777)
Madagascar
Size : 26–39 cm (10–15 in) long[ 115] Habitat : Forest, savanna, shrubland, and grassland[ 116] Diet : Invertebrates, as well as vegetation, fruit, reptiles, amphibians, and small mammals[ 115]
LC
Unknown [ 116]
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^ a b Maree, S. (2015). "Neamblysomus julianae " . IUCN Red List of Threatened Species . 2015 : e.T1089A21285354. doi :10.2305/IUCN.UK.2015-2.RLTS.T1089A21285354.en .
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^ a b c Stephenson, P. J.; Monadjem, A.; Decher, J.; Phalan, B. (2018). "Micropotamogale lamottei " . IUCN Red List of Threatened Species . 2018 : e.T13393A111940150. doi :10.2305/IUCN.UK.2018-2.RLTS.T13393A111940150.en .
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^ Kingdon, Happold, Hoffmann, Butynski, Happold, Kalina , p. 222
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^ a b Garbutt , pp. 40–41
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^ a b c d e f g h i j k l m n o Garbutt , p. 46
^ Garbutt , p. 53
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^ a b Gillespie, Heather (2004). "Microgale dryas " . Animal Diversity Web . University of Michigan . Retrieved November 23, 2022 .
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^ a b Olson, L. E.; Rakotomalala, Z.; Hildebrandt, K. B. P.; Lanier, H. C.; Raxworthy, C. J.; Goodman, S. M. (2009). "Phylogeography of Microgale brevicaudata (Tenrecidae) and Description of a New Species from Western Madagascar" . Journal of Mammalogy . 90 (5): 1095– 1110. doi :10.1644/08-MAMM-A-305.1 .
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Microgale grandidieri " . IUCN Red List of Threatened Species . 2016 : e.T54008309A97190215. doi :10.2305/IUCN.UK.2016-1.RLTS.T54008309A97190215.en .
^ Garbutt , p. 47
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Microgale principula " . IUCN Red List of Threatened Species . 2016 : e.T13350A97201454. doi :10.2305/IUCN.UK.2016-1.RLTS.T13350A97201454.en .
^ Garbutt , p. 56
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Microgale jenkinsae " . IUCN Red List of Threatened Species . 2016 : e.T62015A97192062. doi :10.2305/IUCN.UK.2016-1.RLTS.T62015A97192062.en .
^ Garbutt , p. 52
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Microgale pusilla " . IUCN Red List of Threatened Species . 2016 : e.T41314A97201685. doi :10.2305/IUCN.UK.2016-1.RLTS.T41314A97201685.en .
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Microgale longicaudata " . IUCN Red List of Threatened Species . 2016 : e.T13344A97200213. doi :10.2305/IUCN.UK.2016-1.RLTS.T13344A97200213.en .
^ Stevens, Sarah (2004). "Microgale longicaudata " . Animal Diversity Web . University of Michigan . Retrieved November 23, 2022 .
^ a b Mittermeier, Wilson pp. 170–171
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Microgale majori " . IUCN Red List of Threatened Species . 2016 : e.T62016A97200480. doi :10.2305/IUCN.UK.2016-1.RLTS.T62016A97200480.en .
^ a b Garbutt , p. 54
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Microgale monticola " . IUCN Red List of Threatened Species . 2016 : e.T29462A97200712. doi :10.2305/IUCN.UK.2016-1.RLTS.T29462A97200712.en .
^ a b Trojan, Louis (2022). "Microgale gymnorhyncha " . Animal Diversity Web . University of Michigan . Retrieved November 23, 2022 .
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^ Goodman, S. M.; Soarimalala, V. (2004). "A new species of Microgale (Lipotyphla: Tenrecidae: Oryzorictinae) from the Foret des Mikea of southwestern Madagascar". Proceedings of the Biological Society of Washington . 117 (3): 251– 265.
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Microgale nasoloi " . IUCN Red List of Threatened Species . 2016 : e.T62017A97200965. doi :10.2305/IUCN.UK.2016-1.RLTS.T62017A97200965.en .
^ a b Goodman, S.; Raxworthy, C.; Maminirina, C. P.; Olson, L. (2006). "A new species of shrew tenrec (Microgale jobihely ) from northern Madagascar". Journal of Zoology . 270 (2): 384– 398. doi :10.1111/J.1469-7998.2006.00146.X .
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Microgale jobihely " . IUCN Red List of Threatened Species . 2016 : e.T136628A21282528. doi :10.2305/IUCN.UK.2016-1.RLTS.T136628A21282528.en .
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Microgale fotsifotsy " . IUCN Red List of Threatened Species . 2016 : e.T62013A97191387. doi :10.2305/IUCN.UK.2016-1.RLTS.T62013A97191387.en .
^ Garbutt , p. 51
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Microgale parvula " . IUCN Red List of Threatened Species . 2016 : e.T13349A97201228. doi :10.2305/IUCN.UK.2016-1.RLTS.T13349A97201228.en .
^ a b Mileski, Adam (2004). "Microgale brevicaudata " . Animal Diversity Web . University of Michigan . Retrieved November 23, 2022 .
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^ Garbutt , p. 55
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Microgale soricoides " . IUCN Red List of Threatened Species . 2016 : e.T62018A97201954. doi :10.2305/IUCN.UK.2016-1.RLTS.T62018A97201954.en .
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^ a b Jansa, Sharon (2022). "Microgale dobsoni " . Animal Diversity Web . University of Michigan . Retrieved November 23, 2022 .
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^ a b Belknap, Maureen (2004). "Microgale talazaci " . Animal Diversity Web . University of Michigan . Retrieved November 23, 2022 .
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^ a b Garbutt , pp. 43–44
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Oryzorictes tetradactylus " . IUCN Red List of Threatened Species . 2016 : e.T40591A97203290. doi :10.2305/IUCN.UK.2016-1.RLTS.T40591A97203290.en .
^ a b Fiely, Jonathan (2005). "Oryzorictes hova " . Animal Diversity Web . University of Michigan . Retrieved November 23, 2022 .
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^ a b Garbutt , p. 36
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Echinops telfairi " . IUCN Red List of Threatened Species . 2016 : e.T40592A97188634. doi :10.2305/IUCN.UK.2016-1.RLTS.T40592A97188634.en .
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^ Garbutt , p. 37
^ a b Stephenson, P. J.; Soarimalala, V.; Goodman, S. (2016). "Hemicentetes semispinosus " . IUCN Red List of Threatened Species . 2016 : e.T40593A97189434. doi :10.2305/IUCN.UK.2016-1.RLTS.T40593A97189434.en .
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