2021 in paleomalacology

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This list of fossil molluscs described in 2021 is a list of new taxa of fossil molluscs that were described during the year 2021, as well as other significant discoveries and events related to molluscan paleontology that occurred in 2021.

Ammonites

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Abichites infirmis[1]	Sp. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran
Abichites ovalis[1]	Sp. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran
Agathiceras iranicum[2]	Sp. nov	Valid	Korn & Ghaderi in Korn et al.	Carboniferous-Permian boundary		Iran
Alienisphinctes[3]	Gen. et sp. nov	Valid	Énay & Mangold	Late Jurassic (Kimmeridgian)	Jubaila Formation	Saudi Arabia	A member of the family Ataxioceratidae. The type species is A. cavelieri.
Amaltheus orientalis[4]	Sp. nov	Valid	Nakada et al.	Early Jurassic (Pliensbachian)	Kuruma Group	Japan
Arasella falcata[1]	Sp. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran	An ammonite of uncertain placement, probably a member of the family Xenodiscidae.
Arcestes lawsi[5]	Sp. nov	Valid	Taylor, Guex & Lucas	Late Triassic (Rhaetian)	Gabbs Formation	United States ( Nevada)
Arnioceras ritterbushi[6]	Sp. nov	Valid	Taylor & Guex	Early Jurassic (Sinemurian)	Sunrise Formation	United States ( Nevada)
Arnioceras sparsum[6]	Sp. nov	Valid	Taylor & Guex	Early Jurassic (Sinemurian)	Sunrise Formation	United States ( Nevada)
Artyomites[7]	Gen. et sp. nov	Valid	Smyshlyaeva & Zakharov	Early Triassic (Olenekian)		Russia	A member of the family Palaeophyllitidae. Genus includes new species A. antiquus.
Aulatornoceras frenklerae[8]	Sp. nov	Valid	Söte et al.	Devonian (Frasnian)	Sand Formation	Germany	A member of the family Tornoceratidae.
Aulatornoceras steinhauseni[8]	Sp. nov	Valid	Söte et al.	Devonian (Frasnian)	Sand Formation	Germany	A member of the family Tornoceratidae.
Aulatornoceras ventrosulcatum[8]	Sp. nov	Valid	Söte et al.	Devonian (Frasnian)	Sand Formation	Germany	A member of the family Tornoceratidae.
Boucaulticeras hawthornensis[6]	Sp. nov	Valid	Taylor & Guex	Early Jurassic (Sinemurian)	Sunrise Formation	United States ( Nevada)	A member of the family Schlotheimiidae.
Bullatimorphites szontaghi[9]	Sp. nov	Valid	Galácz
Caloceras avus[10]	Sp. nov	Valid	Kment	Early Jurassic (Hettangian)
Catrisimoceras[11]	Gen. et comb. nov	Valid	Nannarone & Bilotta	Late Jurassic (Tithonian)		Italy	A member of Perisphinctoidea belonging to the family Simoceratidae and possibly to the subfamily Lytogyroceratinae. The type species is "Simoceras" catrianum Zittel (1870).
Cheltonia howarthi[12]	Sp. nov	Valid	Simms & Edmunds	Early Jurassic		United Kingdom
Cibolaites petraschecki[13]	Sp. nov	Valid	Wilmsen et al.	Late Cretaceous (Turonian)	Brießnitz Formation	Germany
Clivotirolites[1]	Gen. et 2 sp. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran	A member of the family Dzhulfitidae. The type species is C. decoratus; genus also includes C. petilus.
Crassotornoceras hetzeneggeri[8]	Sp. nov	Valid	Söte et al.	Devonian (Frasnian)	Sand Formation	Germany	A member of the family Tornoceratidae.
Crassotornoceras nudum[8]	Sp. nov	Valid	Söte et al.	Devonian (Frasnian)	Sand Formation	Germany	A member of the family Tornoceratidae.
Crioceratites claveli[14]	Sp. nov	In press	Bert et al.	Early Cretaceous (Hauterivian)	Vocontian Basin	France
Didymoceras morozumii[15]	Sp. nov	Valid	Misaki & Tsujino	Late Cretaceous (Campanian)	Hiketa Formation	Japan
Dimorphodiscus[16]	Gen. et sp. et comb. nov	Valid	Vermeulen et al.	Early Cretaceous (Barremian)		Algeria Crimea  France	A member of the family Holcodiscidae. The type species is D. lerisae; genus also includes D. uhligi (Karakash, 1907), D. sayni (Joleaud, 1912) and "Metahoplites" postnodosum Vermeulen et al. (2017).
Discoscaphites mullinaxorum[17]	Sp. nov	Valid	Witts et al.	Late Cretaceous (Maastrichtian)	Corsicana Formation	United States ( Texas)
Domanikoceras martenbergense[18]	Sp. nov	Valid	Korn	Devonian (Frasnian)		Germany	A member of the family Tornoceratidae.
Dzhulfites brevisellatus[1]	Sp. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran
Eoasianites baghevangensis[2]	Sp. nov	Valid	Korn & Ghaderi in Korn et al.	Carboniferous-Permian boundary		Iran
Eoermoceras[3]	Gen. et 2 sp. nov	Valid	Énay & Mangold	Middle Jurassic (Bajocian)	Dhruma Formation	Saudi Arabia	A member of the family Stephanoceratidae. The type species is E. glabrum; genus also includes new species E. angulatidomus.
Eolytoceras rohrbachensis[10]	Sp. nov	Valid	Kment	Early Jurassic (Hettangian)
Esfahanites[1]	Gen. et sp. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran	A member of the family Dzhulfitidae. The type species is E. armatus.
Eutomoceras paladeji[19]	Sp. nov	Valid	Tongtherm, Nabhitabhata & Tuanapaya	Triassic	Chaiburi Formation	Thailand	A member of the family Ceratitidae.
Eutomoceras wantanae[19]	Sp. nov	Valid	Tongtherm, Nabhitabhata & Tuanapaya	Triassic	Chaiburi Formation	Thailand	A member of the family Ceratitidae.
Evopharciceras[20]	Gen. et sp. et comb. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Roteisenstein Formation	Germany  Morocco	A member of Agoniatitida belonging to the family Pharciceratidae. The type species is E. formosum; genus also includes "Pharciceras" evolvens Bockwinkel, Becker & Ebbighausen (2009).
Extropharciceras metallicum[20]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Roteisenstein Formation	Germany	A member of Agoniatitida belonging to the family Pharciceratidae.
Galfettites orientalis[7]	Sp. nov	Valid	Smyshlyaeva & Zakharov	Early Triassic (Olenekian)		Russia	A member of the family Galfettitidae.
Glyptophiceras mirum[7]	Sp. nov	Valid	Smyshlyaeva & Zakharov	Early Triassic (Olenekian)		Russia	A member of the family Xenoceltitidae.
Gonioclymenia junggarensis[21]	Sp. nov	In press	Zong, Becker & Ma	Devonian (Famennian)	Hongguleleng Formation	China
Hadrothisbites hanwangensis[22]	Sp. nov	In press	Mietto et al.	Late Triassic (Carnian)	Ma'antang Formation	China
Holcodiscus pseudocaillaudi[16]	Sp. nov	Valid	Vermeulen et al.	Early Cretaceous (Barremian)		France	A member of the family Holcodiscidae.
Hulenites kitamurai[23]	Sp. nov	In press	Matsukawa	Early Cretaceous (Aptian)	Todai Formation	Japan
Inaptychoceras[23]	Gen. et sp. nov	In press	Matsukawa	Early Cretaceous (Aptian)	Todai Formation	Japan	Genus includes new species I. koikei.
Kislovodskoceras[24]	Gen. et sp. nov	Valid	Polkovoy	Early Cretaceous (Aptian)		Russia	A member of the family Desmoceratidae. The type species is K. kislovodskense.
Kurnubiella fallax[3]	Sp. nov	Valid	Énay & Mangold	Middle Jurassic (Callovian)	Upper Dhruma Formation	Saudi Arabia	A member of the family Pachyceratidae.
Laevisimoceras[11]	Gen. et sp. nov	Valid	Nannarone & Bilotta	Late Jurassic (Tithonian)		Italy	A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Parasimoceratinae. The type species is L. inornatum.
Linguatornoceras asbeckense[18]	Sp. nov	Valid	Korn	Devonian (Frasnian)		Germany	A member of the family Tornoceratidae.
Lunupharciceras kochi[20]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Roteisenstein Formation	Germany	A member of Agoniatitida belonging to the family Pharciceratidae.
Lutites[1]	Gen. et 5 sp. et comb. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran	A member of the family Dzhulfitidae. The type species is L. lyriformis; genus also includes new species L. alius, L. paucis, L. plicatus and L. profundus, as well as Alibashites stepanovi Korn & Ghaderi in Korn et al. (2015).
Lytogyroceras tuberculatum[11]	Sp. nov	Valid	Nannarone & Bilotta	Late Jurassic (Tithonian)		Italy	A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Lytogyroceratinae.
Maenioceras ornatum[20]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Roteisenstein Formation	Germany	A member of Agoniatitida belonging to the family Maenioceratidae.
Nejdia duartei[25]	Sp. nov	Valid	Goy & Comas-Rengifo	Early Jurassic (Toarcian)			A member of the family Hildoceratidae.
Obtusicostites gigas[3]	Sp. nov	Valid	Énay & Mangold	Middle Jurassic (Callovian)	Upper Dhruma Formation	Saudi Arabia	A member of the family Perisphinctidae.
Ophimullericeras[26]	Gen. et sp. nov	Valid	Jenks et al.	Early Triassic	Dinwoody Formation	United States ( Nevada)	A member of Ceratitida belonging to the group Meekocerataceae and the family Mullericeratidae. The type species is O. paullae.
Pachyerymnoceras arabicum[3]	Sp. nov	Valid	Énay & Mangold	Middle Jurassic (Callovian)	Tuwaiq Mountain Limestone Formation	Saudi Arabia	A member of the family Pachyceratidae.
Pachyerymnoceras magnum[3]	Sp. nov	Valid	Énay & Mangold	Middle Jurassic (Callovian)	Tuwaiq Mountain Limestone Formation	Saudi Arabia	A member of the family Pachyceratidae.
Palaeophyllites mishkini[7]	Sp. nov	Valid	Smyshlyaeva & Zakharov	Early Triassic (Olenekian)		Russia	A member of the family Palaeophyllitidae.
Paracadoceras vasily[27]	Sp. nov	Valid	Gulyaev in Gulyaev & Ippolitov	Middle Jurassic (Callovian)		Ukraine
Paracochloceras nunminensis[5]	Sp. nov	Valid	Taylor, Guex & Lucas	Late Triassic (Rhaetian)	Gabbs Formation	United States ( Nevada)
Parahoplites inaensis[23]	Sp. nov	In press	Matsukawa	Early Cretaceous (Aptian)	Todai Formation	Japan
Parasimoceras[11]	Gen. et 2 sp. nov	Valid	Nannarone & Bilotta	Late Jurassic (Tithonian)		Italy	A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Parasimoceratinae. The type species is P. venturii; genus also includes P. pseudostenonis.
Paratirolites aduncus[1]	Sp. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran
Paratirolites baghukensis[1]	Sp. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran
Paratirolites lanceolobatus[1]	Sp. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran
Paratirolites robustus[1]	Sp. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran
Paratirolites rubens[1]	Sp. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran
Peripleurites gabbensis[5]	Sp. nov	Valid	Taylor, Guex & Lucas	Late Triassic (Rhaetian)	Gabbs Formation	United States ( Nevada)	A member of Choristocerataceae belonging to the family Rhabdoceratidae.
Pharciceras beyrichi[20]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Roteisenstein Formation	Germany	A member of Agoniatitida belonging to the family Pharciceratidae.
Pharciceras ferrum[20]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Roteisenstein Formation	Germany	A member of Agoniatitida belonging to the family Pharciceratidae.
Pharciceras kruegeri[20]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Roteisenstein Formation	Germany	A member of Agoniatitida belonging to the family Pharciceratidae.
Placites heggi[5]	Sp. nov	Valid	Taylor, Guex & Lucas	Late Triassic (Rhaetian)	Gabbs Formation	United States ( Nevada)	A member of the family Gymnitidae.
Planisimoceras[11]	Gen. et sp. nov	Valid	Nannarone & Bilotta	Late Jurassic (Tithonian)		Italy	A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Parasimoceratinae. The type species is P. costotuberculatum.
Pluripharciceras ahlburgi[20]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Roteisenstein Formation	Germany	A member of Agoniatitida belonging to the family Pharciceratidae.
Pronjaites yolkinensis[28]	Sp. nov	Valid	Mitta	Early Cretaceous		Russia
Pseudogalfettites[7]	Gen. et sp. nov	Valid	Smyshlyaeva & Zakharov	Early Triassic (Olenekian)		Russia	A member of the family Galfettitidae. Genus includes new species P. sakhnoi.
Psiloceras pseudocostosum[10]	Sp. nov	Valid	Kment	Early Jurassic (Hettangian)
Retrotornoceras juxi[8]	Sp. nov	Valid	Söte et al.	Devonian (Frasnian)	Sand Formation	Germany	A member of the family Tornoceratidae.
Rhacophyllites mulleri[5]	Sp. nov	Valid	Taylor, Guex & Lucas	Late Triassic (Rhaetian)	Gabbs Formation	United States ( Nevada)
Rhacophyllites volcanoensis[5]	Sp. nov	Valid	Taylor, Guex & Lucas	Late Triassic (Rhaetian)	Gabbs Formation	United States ( Nevada)
Rollierites biplicatum[29]	Sp. nov	Valid	Jain & Mazur	Middle Jurassic (Callovian)		Poland	A member of the family Pachyceratidae.
Sandbergeroceras archiaci[20]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Roteisenstein Formation	Germany	A member of Agoniatitida belonging to the family Triainoceratidae.
Saudicostites[3]	Gen. et sp. nov	Valid	Énay & Mangold	Middle Jurassic (Bathonian)	Dhruma Formation	Saudi Arabia	A member of Perisphinctoidea of uncertain phylogenetic placement. The type species is S. arabicus.
Shevyrevites corrugatus[1]	Sp. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran	A member of the family Xenodiscidae.
Sigaloceras fundator[27]	Sp. nov	Valid	Gulyaev in Gulyaev & Ippolitov	Middle Jurassic (Callovian)		Ukraine
Sigaloceras kiselevi[27]	Sp. nov	Valid	Gulyaev in Gulyaev & Ippolitov	Middle Jurassic (Callovian)		Ukraine
Simocosticeras[11]	Gen. et sp. nov	Valid	Nannarone & Bilotta	Late Jurassic (Tithonian)		Italy	A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Simoceratinae. The type species is S. biornatum.
Sinotropites[22]	Gen. et sp. nov	In press	Mietto et al.	Late Triassic (Carnian)	Ma'antang Formation	China	Genus includes new species S. sichuanensis.
Sonninia (Euhoploceras) crassicostae[3]	Sp. nov	Valid	Énay & Mangold	Middle Jurassic (Bajocian)	Lower Dhruma Formation	Saudi Arabia	A member of the family Sonniniidae.
Sonninia (Euhoploceras) turbator[3]	Sp. nov	Valid	Énay & Mangold	Middle Jurassic (Bajocian)	Lower Dhruma Formation	Saudi Arabia	A member of the family Sonniniidae.
Sonoraceras[30]	Gen. et sp. nov	In press	Samaniego-Pesqueira, Moreno-Bedmar & Álvarez-Sánchez	Early Cretaceous (Aptian)		Mexico	Genus includes new species S. tepachensis.
Stenopharciceras lotzi[20]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Roteisenstein Formation	Germany	A member of Agoniatitida belonging to the family Pharciceratidae.
Stoliczkaia (Lamnayella) duboisi[31]	Sp. nov	Valid	Kennedy & Latil	Late Cretaceous (Cenomanian)		France	A member of the family Lyelliceratidae.
Stoyanowites parallelus[1]	Sp. nov	Valid	Korn & Hairapetian in Korn et al.	Permian (Changhsingian)	Hambast Formation	Iran	A member of the family Dzhulfitidae.
Strenoceras djangurense[32]	Sp. nov	Valid	Mitta	Middle Jurassic (Bajocian)		Russia
Sulcicostites[11]	Gen. et sp. nov	Valid	Nannarone & Bilotta	Late Jurassic (Tithonian)		Italy	A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Simoceratinae. The type species is S. costatus.
Timorites dendroides[33]	Sp. nov	Valid	Korn & Ghaderi in Tabrizi et al.	Permian (Wuchiapingian)	Julfa Formation	Iran	A member of the family Cyclolobidae.
Tornoceras aequilobum[8]	Sp. nov	Valid	Söte et al.	Devonian (Frasnian)	Sand Formation	Germany	A member of the family Tornoceratidae.
Tornoceras fundibulum[18]	Sp. nov	Valid	Korn	Devonian (Famennian)			A member of the family Tornoceratidae.
Tornoceras occasus[34]	Sp. nov	Valid	Korn	Late Devonian		Morocco	A member of the family Tornoceratidae.
Tornoceras ortus[34]	Sp. nov	Valid	Korn	Late Devonian		Morocco	A member of the family Tornoceratidae.
Tornoceras vretterense[18]	Sp. nov	Valid	Korn	Devonian (Givetian)			A member of the family Tornoceratidae.
Tropaeum evolutum[35]	Sp. nov	In press	Lehmann & Bayliss	Early Cretaceous (Aptian)	Hythe Formation	United Kingdom
Tropidoceras komarovi[36]	Sp. nov	Valid	Zaitsev	Early Jurassic		Crimean Peninsula
Vicininodiceras dalriatense[12]	Sp. nov	Valid	Simms & Edmunds	Early Jurassic		United Kingdom
Volanoceras cuneospinatum[11]	Sp. nov	Valid	Nannarone & Bilotta	Late Jurassic (Tithonian)		Italy	A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Volanoceratinae.
Waehneroceras morawetzi[10]	Sp. nov	Valid	Kment	Early Jurassic (Hettangian)
Wordieoceras mullenae[26]	Sp. nov	Valid	Jenks et al.	Early Triassic	Dinwoody Formation	United States ( Nevada)	A member of Ceratitida belonging to the group Meekocerataceae and the family Ophiceratidae.

Research

• A modern review of the palaeobiology of heteromorph ammonoids is published by Hoffmann et al. (2021), including details of their anatomy, buoyancy, locomotion, predators, diet, palaeoecology, and extinction.[37]
• A study on the impact of the Permian–Triassic extinction event on the morphological diversity of ammonites is published by Dai, Korn & Song (2021).[38]
• A study on the morphology and ontogeny of whorl profiles of the late Anisian ceratitid ammonites is published by Bischof et al. (2021).[39]
• An exceptionally preserved specimen of Sigaloceras enodatum, providing information on the anatomy of ammonite muscles, is described from the Middle Jurassic (Callovian) Kellaways Sand Member of Colne Gravel (Gloucestershire, United Kingdom) by Cherns et al. (2021), who evaluate the implications of this specimen for the knowledge of the swimming mechanism of ammonites.[40]
• Soft parts of a perisphinctid belonging to the genus Subplanites, separated from the conch either taphonomically or during a failed predation, are described from the Tithonian conservation deposits of Wintershof (southern Germany) by Klug et al. (2021).[41]
• The first physical compression experiments on model ammonite septa are performed by Johnson et al. (2021), who attempt to determine whether the increasingly complex fractal margins of ammonite septa might have evolved to increase resistance to shell-crushing predators.[42]
• A study aiming to determine the function of complex septa in the ammonite shell, providing evidence of a relationship between septal complexity and capacity for liquid retention within chambers of the shell, likely improving buoyancy regulation, is published by Peterman et al. (2021).[43]
• Two specimens of Peltoceras athleta having both female and male features are described from Callovian limestone beds in Méron near Montreuil-Bellay (France) by Frau & Boursicot (2021), who interpret these specimens as cases of intersexuality in ammonites.[44]
• Soft tissue remains preserved within the conch of a specimen of Proleymeriella schrammeni from the Albian of Germany are described by Hoffmann et al. (2021).[45]
• A study on the arrangement in space and probable nature of hook-like structures found with specimens of Rhaeboceras halli is published by Smith et al. (2021), who interpret these structures as representing part of the brachial crown armature, and propose a hypothetical reconstruction of an ammonite brachial crown.[46]
• A study on the systematics and phylogeny of the genus Aegocrioceras is published by Weinkauf, Hoffmann & Wiedenroth (2021).[47]
• A study on the hydrostatic properties of Baculites compressus, and on their implications for the knowledge of the swimming capabilities and ecology of this ammonite and orthoconic cephalopods in general, is published by Peterman & Ritterbush (2021).[48]
• Revision of the fossil material of Parapuzosia seppenradensis, including historic specimens from Europe and the United States and new material from England and Mexico, and a study on the ontogeny, distribution and evolution of this ammonite, is published by Ifrim et al. (2021).[49]
• A study on the habitat depth preferences of Hoploscaphites constrictus from the European Boreal Chalk Sea, as indicated by stable isotope thermometry of aptychi and by data from predation marks on the scaphitid moulds, is published by Machalski et al. (2021).[50]
• A study on the relationship between sutural complexity and body size in Cretaceous ammonites is published by Pérez-Claros (2021).[51]

Other cephalopods

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Acrocoelites ippolitae[52]	Sp. nov	Valid	Weis in Weis et al.	Middle Jurassic (Aalenian)		Germany	A belemnite.
Anglonautilus hewaidyi[53]	Sp. nov	In press	Moneer, Bazeen & Kassab	Late Cretaceous (Maastrichtian)		Egypt	A nautiloid cephalopod.
Angulithes monqarensis[53]	Sp. nov	In press	Moneer, Bazeen & Kassab	Late Cretaceous (Maastrichtian)		Egypt	A nautiloid cephalopod.
Arionoceras tokushimaense[54]	Sp. nov	Valid	Niko	Silurian (Ludlow)	Suberidani Group	Japan
Bactroceras fluvii[55]	Sp. nov	Valid	Kröger & Pohle	Ordovician (Dapingian)	Valhallfonna Formation	Norway
Boreopeltis ifrimae[56]	Sp. nov	In press	Fuchs & Stinnesbeck	Late Cretaceous (Turonian)		Mexico
Buttsoceras buldrebreenense[55]	Sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway	A member of the family Troedssonellidae.
Cycloplectoceras hinlopense[55]	Sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway	A member of the family Tarphyceratidae.
Cyclostomiceras profilstrandense[55]	Sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway	A member of the family Cyclostomiceratidae.
Deltoceras beluga[55]	Sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway	A member of the family Cyclostomiceratidae.
Eosomichelinoceras borealis[55]	Sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway	A member of the family Baltoceratidae.
Eosomichelinoceras submedius[57]	Sp. nov	Valid	Evans et al.	Ordovician (Darriwilian)	Pelmis Formation	Iran	A member of the family Baltoceratidae.
Ethanoceras[55]	Gen. et sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway	A relative of Rioceras. The type species is E. solitudines.
Gzheloceras aisenvergi[58]	Sp. nov	Valid	Dernov	Carboniferous (Serpukhovian)	Samara Formation	Ukraine	A member of the family Tainoceratidae.
Hemichoanella occulta[55]	Sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway
Hinlopoceras[55]	Gen. et 2 sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway	A relative of Rioceras. The type species is H. tempestatis; genus also includes H. venti.
Jovellania praetermissa[59]	Sp. nov	Valid	Brauckmann & Koch	Devonian (Givetian)	Massenkalk Formation	Germany
Knightoceras extorris[58]	Sp. nov	Valid	Dernov	Carboniferous (Pennsylvanian)		Ukraine	A member of the family Koninckioceratidae.
Lawrenceoceras ebenus[55]	Sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway
Lawrenceoceras larus[55]	Sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway
Litoceras profilbekkenense[55]	Sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway
Mojsvaroceras gianii[60]	Sp. nov	Valid	Pieroni & Prinoth	Middle Triassic (Anisian)		Italy	A nautiloid.
Nyfrieslandoceras[55]	Gen. et sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway	A member of Oncocerida belonging to the family Phthanoncoceratidae. The type species is N. bassleroceroides.
Olenidslettoceras[55]	Gen. et sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway	A member of Oncocerida belonging to the family Phthanoncoceratidae. The type species is O. farmi.
Paracenoceras waageni[61]	Sp. nov	Valid	Halder	Middle Jurassic (Callovian)	Chari Formation	India	A member of the family Paracenoceratidae.
Planetoceras yefimenkoi[58]	Sp. nov	Valid	Dernov	Carboniferous (Bashkirian)	Mospinka Formation	Ukraine	A member of the family Koninckioceratidae.
Proterocameroceras valhallfonnense[55]	Sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway
Protocycloceras minor[55]	Sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway	A member of the family Troedssonellidae.
Rhynchorthoceras yizanense[62]	Sp. nov	Valid	Fang in Fang et al.	Ordovician		China	A member of Lituitida.
Sinoceras complexum[62]	Sp. nov	Valid	Fang in Fang et al.	Ordovician		China	A member of Lituitida.
Sorosoceras[57]	Gen. et sp. nov	Valid	Evans et al.	Ordovician (Darriwilian)	Pelmis Formation	Iran	A member of the family Geisonoceratidae. The type species is S. castellum.
Svalbardoceras[55]	Gen. et 2 sp. nov	Valid	Kröger & Pohle	Ordovician (Floian)	Valhallfonna Formation	Norway	A relative of Rioceras. The type species is S. sterna; genus also includes S. skua.
Tyrioceras longicameratum[62]	Sp. nov	Valid	Fang in Fang et al.	Ordovician		China	A member of Lituitida.
Xiaohenautilus mulleni[26]	Sp. nov	Valid	Jenks et al.	Early Triassic	Dinwoody Formation	United States ( Nevada)	A member of Nautilida belonging to the group Trigonoceratoidea and the family Grypoceratidae.

Research

• New fossil material with shell features that characterize early cephalopods is described from the Cambrian (Terreneuvian) Bonavista Formation (Canada) by Hildenbrand et al. (2021), who interpret this finding as possibly representing the earliest cephalopods known to date, potentially pushing the origin of cephalopods back in time by about 30 million years, to the time before the first occurrence of euarthropods.[63]
• Two specimens of Syrionautilus libanoticus preserving soft parts, including a specimen showing an imprint of the conch, are described from the Cenomanian of Hadjoula (Lebanon) by Klug et al. (2021).[64]
• A study on the diversity patterns and spatial structure of belemnite assemblages in Western Tethys Ocean during the Early Jurassic is published by Neige, Weis & Fara (2021).[65]
• A study on morphological changes of belemnites across the Pliensbachian-Toarcian boundary event, based on data from the Peniche GSSP section (Portugal), is published by Nätscher et al. (2021).[66]
• A study on chemistry, organization and genesis of circular structures (superficially resembling chromatophores) preserved in coleoid cephalopod specimens from the Jurassic of Germany and the Cretaceous of Lebanon is published by Klug et al. (2021).[67]
• A well-preserved soft-body imprint of a fossil squid is described from the lower Oligocene of the Krasnodar region (Russia) by Mironenko et al. (2021), who interpret this finding as the first unquestionable representative of Teuthida in the fossil record known to date, and report preservation of fish remains in the stomach contents of this squid, indicative of its piscivorous diet.[68]
• A specimen of Jeletzkyteuthis coriacea holding a specimen of Parabelopeltis flexuosa in its arms, with the apex very close to its jaws, is described from the Toarcian Posidonia Shale (Germany) by Klug et al. (2021), who interpret this finding as a case of frozen predatory behaviour of two different octobrachian species.[69]
• Redescription of the gladius of the holotype specimen of Necroteuthis hungarica is published by Košťák et al. (2021), who reinterpret this species as a member of the family Vampyroteuthidae linking Mesozoic loligosepiids with extant Vampyroteuthis.[70]
• Klompmaker & Landman (2021) describe the oldest drill holes produced by octopodoids in lucinid bivalve specimens from the Campanian of South Dakota, indicating that the drilling habit evolved early in the evolutionary history of Octopodoidea.[71]

Gastropods

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Acirsa ambrosii[72]	Sp. nov	Valid	Brunetti & Cresti	Pliocene		Italy	A species of Acirsa.
Acirsa (Notacirsa) texana[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Acirsa.
Acteon pseudotypica[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Acteon.
Acutitomaria[74]	Gen. et sp. nov	Valid	Karapunar & Nützel	Late Triassic	San Cassiano Formation	Italy	A member of Pleurotomariida. Genus includes new species A. kustatscherae.
Alatostrombus[75]	Gen. et sp. et comb. nov	Valid	Raven	Miocene		Indonesia  Malaysia	A member of the family Strombidae. Genus includes new species A. acutalatus, as well as "Strombus" sedanensis Martin (1899).
Alvania absuturalis[76]	Sp. nov	Valid	Chirli & Forli	Pliocene		Italy	A species of Alvania.
Alvania ameliae[76]	Sp. nov	Valid	Chirli & Forli	Pleistocene		Italy	A species of Alvania.
Alvania benestarensis[76]	Sp. nov	Valid	Chirli & Forli	Miocene		Italy	A species of Alvania.
Alvania cerreti[77]	Sp. nov	Valid	Gardella et al.	Pliocene		Italy	A species of Alvania.
Alvania pluricosticillata[76]	Sp. nov	Disputed	Chirli & Forli	Plio-Pleistocene		Italy	Originally described as a species of Alvania. Argued to be a junior synonym of the extant taxon Crisilla gagliniae by Amati & Oliverio (2024).[78]
Amaea (Confusiscala) paleocaenica[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Ammonicera danieli[79]	Sp. nov	Valid	Chernyshev & Goedert	Late Eocene/early Oligocene	Gries Ranch Formation	United States ( Washington)	A species of Ammonicera.
Ammonicera rolani[79]	Sp. nov	Valid	Chernyshev & Goedert	Late Eocene/early Oligocene	Gries Ranch Formation	United States ( Washington)	A species of Ammonicera.
Ampezzogyra angulata[80]	Sp. nov	Valid	Nützel & Hausmann in Hausmann et al.	Late Triassic (Carnian)	San Cassiano Formation	Italy	A member of the family Stuoraxidae.
Amphidromus ubaldii[81]	Sp. nov	Valid	Dharma	Late Pliocene		Indonesia	A species of Amphidromus.
Amplitomaria[74]	Gen. et comb. nov	Valid	Karapunar & Nützel				A member of Pleurotomariida. The type species is "Pleurotomaria" spuria Münster (1841).
Anacithara akisi[82]	Sp. nov	Valid	Kolokotronis	Pliocene		Cyprus	A species of Anacithara.
Anceps siminescui[83]	Nom. nov	Valid	Harzhauser	Middle-late Miocene		Moldova	A Trochidae member; a replacement name for Trochus semistriatus Siminescu & Barbu (1940).
Angulatella[80]	Gen. et sp. nov	Valid	Nützel & Hausmann in Hausmann et al.	Late Triassic (Carnian)	San Cassiano Formation	Italy	A member of the family Prostyliferidae. The type species is A. bizzarinii.
Antithala[84]	Gen. et comb. et 2 sp. nov	Valid	Harzhauser & Landau	Miocene	Dej Formation	Bosnia and Herzegovina  Hungary  Poland  Romania  Ukraine	A member of the family Costellariidae. The type species is "Mitra" sturi Hoernes & Auinger (1880); genus also includes "Mitra" neugeboreni Hoernes & Auinger (1880), as well as new species A. claviformis and A. filipescui.
Apiocypraea cerretensis[85]	Sp. nov	Valid	Fehse et al.	Pliocene (Zanclean)		Italy	A member of the family Eocypraeidae.
Aplus abessensis[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Aplus.
Aplus grandiculus[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Aplus.
Aplus littoralis[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Aplus.
Aplus pseudoaquitanensis[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Aplus.
Aquilofusus? antiqua[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Ascheria canni[87]	Sp. nov	Valid	Kaim et al.	Late Cretaceous (Cenomanian to Turonian)		Cyprus	A member of the family Hokkaidoconchidae.
Asiaharpa[88]	Gen. et sp. nov	Valid	Raven	Miocene		Malaysia	A member of the family Harpidae. Genus includes new species A. sarawakiana.
Attenuata? danica[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Possibly a species of Attenuata.
Auingeria[89]	Gen. et comb. nov	Valid	Harzhauser & Landau	Miocene	Baden Formation	Austria  Czech Republic  Germany  Hungary	A member of the family Columbellidae. The type species is "Columbella" bellardii Hörnes (1852); genus also includes "Columbella (Anachis)" austriaca Hoernes & Auinger (1880) and "Pyrene (Anachis) bellardii" grussbachensis Csepreghy-Meznerics (1969; raised to the rank of a separate species Auingeria grussbachensis).
Axonella[90]	Gen. et comb. et sp. nov	Valid	Pacaud, Lhomme & Renaud	Eocene (Ypresian)		France  Spain	A member of the family Hemisinidae. The type species is "Cerithium" palense Rouault (1850); genus also includes "Potamides" granosus Doncieux (1908), as well as new species A. falsiacum.
Babylonia chattica[91]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Babylonia.
Babylonia edigittenbergeri[92]	Sp. nov	Valid	Raven	Miocene	Miri Formation	Malaysia	A species of Babylonia.
Babylonia sarawakiana[92]	Sp. nov	Valid	Raven	Miocene	Miri Formation	Malaysia	A species of Babylonia.
Bandellina compacta[80]	Sp. nov	Valid	Nützel & Hausmann in Hausmann et al.	Late Triassic (Carnian)	San Cassiano Formation	Italy	A member of the family Cornirostridae.
Barycypraea abbasi[93]	Sp. nov	Valid	Dovesi & Parsons	Miocene (Serravallian)	Nyalindung Formation	Indonesia	A species of Barycypraea.
Barycypraea alessandrodovesi[93]	Sp. nov	Valid	Dovesi & Parsons	Miocene (Serravallian)	Nyalindung Formation	Indonesia	A species of Barycypraea.
Bela pediensis[94]	Sp. nov	Valid	Kolokotronis	Pliocene		Cyprus	A species of Bela.
Bellacolumbella[89]	Gen. et comb. nov	Valid	Harzhauser & Landau	Miocene	Baden Formation	Austria  Czech Republic  Hungary  Romania	A member of the family Columbellidae. The type species is "Columbella (Nitidella)" karreri Hoernes & Auinger (1880); genus also includes "Columbella (Nitidella)" embryonalis Boettger (1902) and "Columbella (Nitidella)" katharinae Hoernes & Auinger (1880).
Bellardithala[84]	Nom. et 4 sp. nov	Valid	Harzhauser & Landau	Miocene and Pliocene	Baden Formation Dej Formation	Austria  Czech Republic  Italy  Poland  Romania  Ukraine  Hungary?	A member of the family Costellariidae; a replacement name for Micromitra Bellardi (1888). The type species is "Voluta" obsoleta Brocchi (1814); genus also includes new species B. kovaci, B. baluki, B. fedosovi and B. dacica.
Bieleraxis[73]	Gen. et 2 sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Genus includes new species B. discoidea and B. altis.
Bithynia disregularis[95]	Sp. nov	In press	Li et al.	Oligocene	Yehucheng Formation	China	A species of Bithynia.
Bithynia obliquus[95]	Sp. nov	In press	Li et al.	Oligocene	Yehucheng Formation	China	A species of Bithynia.
Bithynia parainclinimarga[95]	Sp. nov	In press	Li et al.	Oligocene	Yehucheng Formation	China	A species of Bithynia.
Bithynia paramonolithic[95]	Sp. nov	In press	Li et al.	Oligocene	Yehucheng Formation	China	A species of Bithynia.
Bittium prekoeneni[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Bittium.
Bulimulus frenguellii[96]	Sp. nov	Valid	Cabrera et al.	Late Cretaceous	Queguay Formation	Uruguay	A species of Bulimulus.
Burminella[97]	Gen. et sp. nov	In press	Balashov, Perkovsky & Vasilenko	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A land snail possessing morphological traits intermediate between members of the family Pupinidae and other members of Cyclophoroidea. Genus includes new species B. artiukhini.
Bursa transeuntis[98]	Sp. nov	Valid	Forli & Smriglio	Miocene and Pliocene		Hungary  Italy  Poland	A species of Bursa.
Caecum texanum[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Caecum.
Califrapana[99]	Gen. et comb. nov	Valid	Powell & Houart	Late Oligocene to early Miocene	Vaqueros Formation	Mexico  United States ( California)	A member of the family Muricidae. The type species is "Purpura" vaquerosensis Arnold (1907).
Calyptraphorus velusus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Camponaxis bandeli[100]	Sp. nov	Valid	Pieroni, Monari & Todd	Late Triassic (Carnian)	San Cassiano Formation	Italy	A Tofanellidae member
Campylacrum texanum[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Canariella (Majorata) gerti[101]	Sp. nov	Valid	Groh	Quaternary		Canary Islands	A species of Canariella.
Cancellotomaria[74]	Gen. et comb. nov	Valid	Karapunar & Nützel				A member of Pleurotomariida. The type species is "Pleurotomaria" subcancellata d'Orbigny (1850).
Cancilla nanostriatula[102]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Baden Formation	Austria  Bulgaria  Hungary	A species of Cancilla.
Cancilla wagreichi[102]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Dej Formation	Romania	A species of Cancilla.
Capulus gelus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Capulus.
Cassis kalmani[103]	Sp. nov	Valid	Vicián & Kovács	Eocene (Lutetian)	Csernye Formation	Hungary	A species of Cassis.
Cassis rasae[104]	Sp. nov	Valid	Petuch & Berschauer	Pleistocene		United States ( Florida)	A species of Cassis.
Cassis viliusi[104]	Sp. nov	Valid	Petuch & Berschauer	Pleistocene		United States ( Florida)	A species of Cassis.
Cenorhytis[73]	Gen. et sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Genus includes new species C. groesbecki.
Ceratopea? moridunensis[105]	Sp. nov	Valid	Ebbestad & Cope	Ordovician (Floian)		United Kingdom	A member of the family Raphistomatidae.
Cerithiella acuta[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Cerithiella.
Cerithiscala parvibrazosa[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Cerithiscala tehuacana[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Chemnitzia silvai[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Chemnitzia.
Chevallieria texanopsis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Chicomurex parvus[107]	Sp. nov	Valid	Merle, Landau & Breitenberger	Miocene		Indonesia	A species of Chicomurex.
Chicoreus javanus[107]	Sp. nov	Valid	Merle, Landau & Breitenberger	Miocene		Indonesia	A species of Chicoreus.
Chicoreus martini[107]	Sp. nov	Valid	Merle, Landau & Breitenberger	Miocene		Indonesia	A species of Chicoreus.
Chicoreus solidus[107]	Sp. nov	Valid	Merle, Landau & Breitenberger	Miocene		Indonesia	A species of Chicoreus.
Clava (Semivertagus) scriba[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Clavatula hirmetzli[108]	Sp. nov	Valid	Kovács & Vicián	Middle Miocene	Lajta Limestone Formation	Hungary	A species of Clavatula.
Clavatula santhai[108]	Sp. nov	Valid	Kovács & Vicián	Middle Miocene	Lajta Limestone Formation	Hungary	A species of Clavatula.
Clavatula szekelyhidiae[108]	Sp. nov	Valid	Kovács & Vicián	Middle Miocene	Lajta Limestone Formation	Hungary	A species of Clavatula.
Cochlespira (Tahusyrinx) laevis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Cochlespira.
Conilithes herodus[109]	Sp. nov	Valid	Psarras, Koskeridou & Merle	Miocene (Tortonian)		Greece	A species of Conilithes.
Conradconfusus nodulinus[110]	Sp. nov	Valid	Garvie	Eocene	Cook Mountain Formation	United States ( Texas)
Conradconfusus texanus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Conus (Kalloconus) asterousiaensis[109]	Sp. nov	Valid	Psarras, Koskeridou & Merle	Miocene (Tortonian)		Greece  Italy	A species of Conus.
Conus (Kalloconus) helladicus[109]	Sp. nov	Valid	Psarras, Koskeridou & Merle	Miocene (Tortonian)		Greece	A species of Conus.
Cordieria biouesensis[111]	Sp. nov	Valid	Pacaud	Paleocene (Thanetian)		France	A species of Cordieria.
Coronia argilla[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Costoanachis venzoi[89]	Nom. nov	Valid	Harzhauser & Landau	Miocene		Italy	A species of Costoanachis; a replacement name for Anachis (Costoanachis) subcorrugata Venzo & Pelosio (1963).
Crassaseila[73]	Gen. et 2 sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Genus includes new species C. ripleyana and C. paleocenica.
Crassispira plioibericostricta[112]	Sp. nov	Valid	Brunetti & Forli	Pliocene (Zanclean)		Spain	A species of Crassispira.
Crassispira pseudoglabra[112]	Sp. nov	Valid	Brunetti & Forli	Pliocene (Piacenzian)		Italy	A species of Crassispira.
Crenaturricula minuta[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Creonella hebetertia[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Cretatortulosa cretakachinensis[113]	Sp. nov	In press	Yu, Salvador & Jarzembowski	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar
Cretatortulosa gignens[114][115][116]	Sp. nov	In press	Jochum, Yu & Neubauer	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Pupinidae.
Cryptochorda paleocaenica[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Cyathopoma pingyiensis[117]	Sp. nov	In press	Yu et al.	Cretaceous–Paleogene transition	Pingyi Basin	China	A species of Cyathopoma.
Cylichna (Cylichnopsis) elliptica[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Cylichna.
Cymatiella dulaii[103]	Sp. nov	Valid	Vicián & Kovács	Eocene (Lutetian)	Csernye Formation	Hungary	A species of Cymatiella.
Cymatiella tzankovi[103]	Nom. nov	Valid	Vicián & Kovács	Paleocene		Bulgaria	A species of Cymatiella; a replacement name for Eutritonium (Sassia) rutoti Tzankov (1940).
Cymatium danianum[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Cymatium.
Cyphoma guraboensis[118]	Sp. nov	Valid	Groves & Landau	Early Pliocene	Gurabo Formation	Dominican Republic	A species of Cyphoma.
Cypraeorbis alisonkayae[118]	Sp. nov	Valid	Groves & Landau	Miocene		Dominican Republic	A species of Cypraeorbis.
Cyprioconcha[87]	Gen. et sp. nov	Valid	Kaim et al.	Late Cretaceous (Cenomanian to Turonian)		Cyprus	A member of the family Hokkaidoconchidae. The type species is C. robertsoni.
Defensina[89]	Gen. et comb. nov	Valid	Harzhauser & Landau	Miocene		Bosnia and Herzegovina  Bulgaria  Romania	A member of the family Columbellidae. The type species is "Columbella (Mitrella)" bucciniformis Hoernes & Auinger (1880).
Dermomurex wanneri[107]	Nom. nov	Valid	Merle, Landau & Breitenberger	Miocene		Indonesia	A species of Dermomurex; a replacement name for Murex acuticostatus Wanner & Hahn (1935).
Desbruyeresia kambiaensis[87]	Sp. nov	Valid	Kaim et al.	Late Cretaceous (Cenomanian to Turonian)		Cyprus	A species of Desbruyeresia.
Desbruyeresia kinousaensis[87]	Sp. nov	Valid	Kaim et al.	Late Cretaceous (Cenomanian to Turonian)		Cyprus	A species of Desbruyeresia.
Desbruyeresia memiensis[87]	Sp. nov	Valid	Kaim et al.	Late Cretaceous (Cenomanian to Turonian)		Cyprus	A species of Desbruyeresia.
Discotectonica paracutissima[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Discotectonica.
Doliella parnitens[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Doliella.
Domiporta amoena[102]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Dej Formation	Austria  Bulgaria  Romania	A species of Domiporta.
Domiporta pulchra[102]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Dej Formation	Romania	A species of Domiporta.
Domiporta turpis[102]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Baden Formation	Austria	A species of Domiporta.
Ederazyga[100]	Gen. et sp. et comb. nov	Valid	Pieroni, Monari & Todd	Late Triassic (Carnian to Rhaetian)	Nayband Formation San Cassiano Formation Zu Limestone	Iran  Italy  Slovenia	A possible Zygopleuridae member. The type species is E. fanchini; genus also includes "Cerithium"? lateplicatum Klipstein (1843).
Ellobium mizutanii[119]	Sp. nov	Valid	Kawase & Ichihara	Early Miocene	Akeyo Formation	Japan	A species of Ellobium.
Entemnotrochus kathiawarensis[120]	Sp. nov	Valid	Bose, Das & Mondal	Miocene (Burdigalian)		India	A species of Entemnotrochus.
Eoacteon paracutissima[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Eoancilla lapicidina[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Eocithara (?) laurencei[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Eopleurotoma atacica[111]	Nom. nov	Valid	Pacaud	Eocene (Ypresian)		France	A species of Eopleurotoma; a replacement name for Pleurotoma (Eopleurotoma) romani Doncieux (1908).
Eotrivia pacaudi[121]	Sp. nov	Valid	Fehse	Eocene		Italy
Eoturris fallsensis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Eoworthenia frydai[74]	Nom. nov	Valid	Karapunar & Nützel				A member of Pleurotomariida; a replacement name for Worthenia rarissima Barrande.
Episcomitra antibellardii[102]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Dej Formation	Romania	A species of Episcomitra.
Episcomitra leopoldiana[102]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Baden Formation	Austria  Czech Republic  Poland	A species of Episcomitra.
Episcomitra missile[102]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Grund Formation	Austria  Hungary	A species of Episcomitra.
Episcomitra neubaueri[102]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Dej Formation	Romania	A species of Episcomitra.
Episcomitra pontonxensis[91]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Episcomitra.
Episcomitra pseudoincognita[102]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Dej Formation	Romania  Ukraine	A species of Episcomitra.
Episcomitra vermeuleni[122]	Sp. nov	Valid	Pacaud	Eocene (Priabonian)		Ukraine	A species of Episcomitra.
Eulimella ariejansseni[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Eulimella.
Euthema myanmarica[123]	Sp. nov	Valid	Balashov	Cretaceous	Hkamti amber	Myanmar	A species of Euthema.
Euthria mellianensis[86]	Sp. nov	Valid	Lozouet	Miocene		France	A species of Euthria.
Euthria tarusatensis[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Euthria.
Euthriofusus argillensis[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France
Falsifusus problematicus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Fedosovia[84]	Gen. et comb. nov	Valid	Harzhauser & Landau	Miocene		Czech Republic  Italy  Romania	A member of the family Costellariidae. The type species is "Mitra (Callithea)" fuchsi Hoernes & Auinger (1880).
Fraudiziba[102]	Gen. et nom. et sp. nov	Valid	Harzhauser & Landau	Miocene and Pliocene	Karaman Basin	Austria  Bosnia and Herzegovina  Czech Republic  Hungary  Italy  Poland  Romania  Slovakia  Turkey  Ukraine	A member of the family Mitridae. The type species is F. paratethyca (a replacement name for Mitra austriaca Csepreghy-Meznerics, 1950); genus also includes new species F. ottomanica.
Fulgurofusus spinosus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Fulgurofusus.
Fusulculus texanus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Fusulculus.
Galba prima[124]	Sp. nov	Valid	Yu, Neubauer & Jochum	Cretaceous	Burmese amber	Myanmar	A species of Galba.
Gemmula bearrizensis[111]	Nom. nov	Valid	Pacaud	Eocene (Priabonian)		France	A species of Gemmula; a replacement name for Drillia pellati Boussac (1911).
Gemmula osca[111]	Sp. nov	Valid	Pacaud	Eocene (Bartonian)		Spain	A species of Gemmula.
Gentilifulgur[73]	Gen. et sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Genus includes new species G. praemortoni.
Gibbula lovellreevei[83]	Nom. nov	Valid	Harzhauser	Pliocene		United Kingdom	A Trochidae member; a replacement name for Trochus (Gibbula) reevei Harmer (1923).
Gibbula pizzolatoi[125]	Sp. nov	Valid	Spadini	Pliocene		Italy	A species of Gibbula.
Gibbula steiningeri[83]	Nom. nov	Valid	Harzhauser	Early Miocene		Austria	A Trochidae member; a replacement name for Trochus amedei bicincta Schaffer (1912).
Gibbula tavanii[83]	Nom. nov	Valid	Harzhauser	Miocene		Libya	A Trochidae member; a replacement name for Gibbula minima Tavani (1939).
Gibbuliculus[83]	Gen. et comb. et nom. nov	Valid	Harzhauser	Oligocene to Pleistocene		Europe	A Trochidae member; The type species is "Gibbula" brebioni Landau, Van Dingenen & Ceulemans (2017). Genus includes G. saccoi (a replacement name for Gibbula protumida Sacco, 1896).
"Gibbulinella" sandbergeri[126]	Nom. nov	Valid	Harzhauser & Neubauer	Eocene		Italy	A member of the family Streptaxidae; a replacement name for Pupa simplex Sandberger (1870).
Haedropleura dellabellai[72]	Sp. nov	Valid	Brunetti & Cresti	Pliocene		Italy	A species of Haedropleura.
Hammatospira cancellata[127]	Sp. nov	Valid	Mazaev	Early Carboniferous		Kazakhstan
Hespererato canae[118]	Sp. nov	Valid	Groves & Landau	Late Miocene and early Pliocene	Cercado Formation	Dominican Republic	A species of Hespererato.
Hespererato praeclusa[118]	Sp. nov	Valid	Groves & Landau	Early–middle Miocene	Baitoa Formation	Dominican Republic	A species of Hespererato.
Hexaplex nascium[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Hexaplex.
Hirsuticyclus canaliculatus[128]	Sp. nov	In press	Yu	Cretaceous	Burmese amber	Myanmar	A member of the family Cyclophoridae.
Hochheimia[126]	Nom. nov	Valid	Harzhauser & Neubauer	Oligocene		Germany	A member of the family Hygromiidae; a replacement name for Palaeotrichia Nordsieck (2014).
Hokkaidoconcha morisseaui[87]	Sp. nov	Valid	Kaim et al.	Late Cretaceous (Cenomanian to Turonian)		Cyprus	A member of the family Hokkaidoconchidae.
Holospira? fallsensis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Possibly a species of Holospira.
?Hydrocena praecursor[129]	Sp. nov	In press	Yu & Neubauer	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Hydrocenidae.
Jponsia lacetania[130]	Sp. nov	Valid	Pacaud & Goret	Eocene (Ypresian)		Spain	A member of the family Pachychilidae.
Kapalmerella gelus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Keepingia elongata[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France
Laevispira[75]	Gen. et comb. et sp. nov	Valid	Raven	Miocene and Pliocene		Brunei  India  Indonesia	A member of the family Strombidae. Genus includes "Dolomena" bruneiensis Harzhauser, Raven & Landau (2018), "Strombus" glaber Martin (1879), "Strombus" karikalensis Cossmann (1903), "Strombus" triangulatus Martin (1879) and "Strombus" varinginensis Martin (1899), as well as new species L. seriaensis.
Laevitrivia[131]	Gen. et comb. et 5 sp. nov	Valid	Fehse & Vicián	Miocene and Pliocene		France  Italy	A member of the family Triviidae. The type species is "Cypraea" pisolina Lamarck, (1810); genus also includes new species L. annae, L. bertaccinii, L. pliopisolina, L. pseudocalva and L. ventricostata.
Latirus? asperpellis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Possibly a species of Latirus.
Laubella subsulcata[74]	Sp. nov	Valid	Karapunar & Nützel	Late Triassic	San Cassiano Formation	Italy	A member of Pleurotomariida.
Lineacingulum[74]	Gen. et comb. nov	Valid	Karapunar & Nützel				A member of Pleurotomariida. The type species is "Pleurotomaria" texturata Münster (1841).
Lineaetomaria[74]	Gen. et comb. nov	Valid	Karapunar & Nützel				A member of Pleurotomariida. The type species is "Pleurotomaria" decorata Münster (1841).
Liostomia wilvanderstoelae[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Liostomia.
Littorina elongata[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Littorina.
Littorina (Prosthenodon) incertus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Littorina.
Lupira danianum[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Lutema gracilis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Lyria elongata[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Lyria.
Lyria tehuacana[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Lyria.
Malea hyaducki[104]	Sp. nov	Valid	Petuch & Berschauer	Pliocene		United States ( Florida)	A species of Malea.
Martaia[89]	Gen. et comb. nov	Valid	Harzhauser & Landau	Miocene		Austria  Bulgaria  Czech Republic  Hungary  Romania  Ukraine	A member of the family Columbellidae. The type species is "Columbella (Anachis)" dujardini Hoernes & Auinger (1880); genus also includes "Columbella (Anachis)" zitteli Hoernes & Auinger (1880).
Mathilda (Echinimathilda) duogenta[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Mathilda.
Melongena pontonxensis[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Melongena.
Metula bulbiformis[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Metula.
Metula nana[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Metula.
Metula pseudomajor[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Metula.
Metula rivulisensis[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Metula.
Mimospira llangynogensis[105]	Sp. nov	Valid	Ebbestad & Cope	Ordovician (Floian)		United Kingdom	A member of the family Clisospiridae.
Mitrella dacica[89]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Dej Formation	Hungary  Romania  Slovakia	A species of Mitrella.
Mitrella demaintenonae[89]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Baden Formation Dej Formation	Austria  Romania	A species of Mitrella.
Mitrella elongatissima[89]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Baden Formation	Austria  Bosnia and Herzegovina	A species of Mitrella.
Mitrella fortis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Mitrella.
Mitrella tarbelliana[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Mitrella.
Mitrella viennensis[89]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Baden Formation	Austria  Bulgaria  Czech Republic  Kazakhstan  Poland  Romania	A species of Mitrella.
Moniquia astibiai[130]	Sp. nov	Valid	Pacaud & Goret	Eocene (Priabonian)		Spain	A member of the family Pachychilidae.
Monoplex s.l. szakonyii[103]	Sp. nov	Valid	Vicián & Kovács	Eocene (Lutetian)	Csernye Formation	Hungary	A member of the family Cymatiidae.
Mulderia mulderi[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A member of the family Pyramidellidae.
Multitrochia[73]	Gen. et sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Genus includes new species M. tehuacana.
Nannamoria alquezae[132]	Sp. nov	Valid	Hawke	Miocene		Australia	A species of Nannamoria.
Nannamoria cadella[132]	Sp. nov	Valid	Hawke	Miocene	Cadel Formation	Australia	A species of Nannamoria.
Nannamoria costatum[132]	Sp. nov	Valid	Hawke	Pliocene	Cameron Inlet Formation	Australia	A species of Nannamoria.
Nannamoria flindersi[132]	Sp. nov	Valid	Hawke	Pliocene	Cameron Inlet Formation	Australia	A species of Nannamoria.
Nannamoria gnotuki[132]	Sp. nov	Valid	Hawke			Australia	A species of Nannamoria.
Nannamoria hiscocki[132]	Sp. nov	Valid	Hawke	Pliocene	Cameron Inlet Formation	Australia	A species of Nannamoria.
Nannamoria malonei[132]	Sp. nov	Valid	Hawke			Australia	A species of Nannamoria.
Nannamoria persimilis[132]	Sp. nov	Valid	Hawke	Miocene		Australia	A species of Nannamoria.
Naria praehelvola[133]	Sp. nov	Valid	Fehse & Vicián	Miocene		Indonesia	A species of Naria.
Natica (Carinacca) kincaidensis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Natica.
Nebularia soliphila[102]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Baden Formation	Austria	A species of Nebularia.
Nemaspira[134]	Gen. et comb. et 2 sp. nov	Valid	Mazaev	Early Carboniferous to Late Triassic		Belgium  Russia  Thailand  United Kingdom  United States  China?	A member of the family Gosseletinidae. Genus includes several species formerly assigned to the genera Gosseletina and Globodoma, as well as new species N. shamaevi and N. skuini.
Nitidiclavus senesi[135]	Sp. nov	Valid	Biskupič	Miocene		Slovakia	A member of the family Drilliidae.
Niveria callosa[136]	Sp. nov	Valid	Fehse	Late Oligocene		France	A species of Niveria.
Niveria novosperara[136]	Sp. nov	Valid	Fehse	Late Oligocene		France	A species of Niveria.
Nodocingulum[74]	Gen. et comb. et 2 sp. nov	Valid	Karapunar & Nützel	Late Triassic	San Cassiano Formation	Italy	A member of Pleurotomariida. The type species is "Pleurotomaria" coronata Münster (1841); genus also includes new species N. ernstkittli and possibly also N? turris.
Nodolargena[73]	Gen. et sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A member of the new family Nodolargenidae. Genus includes new species N. frosti.
Nodovoluta[73]	Gen. et sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Genus includes new species N. coronata.
Odostomia malagensis[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Odostomia.
Oligonalia[86]	Gen. et comb. nov	Valid	Lozouet	Oligocene		France	A new genus for "Siphonalia" riparia Lozouet (1999).
Olinderia[91]	Gen. et comb. et 3 sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A member of the family Charitodoronidae. The type species is "Charitodoron" tauzini Lozouet (1991); genus also includes new species O. lestrillensis, O. poustagnacensis and O. verdunensis.
Olivella tehuacanensis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Olivella.
Ondina pinguis[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Ondina.
Onustus undosus[75]	Sp. nov	Valid	Raven	Miocene		Brunei	A species of Onustus.
Opalia tehuacana[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Opalia.
Orbitestella kieli[79]	Sp. nov	Valid	Chernyshev & Goedert	Late Eocene/early Oligocene	Gries Ranch Formation	United States ( Washington)	A member of the family Orbitestellidae.
Orthosurcula ethani[110]	Sp. nov	Valid	Garvie	Eocene	Cook Mountain Formation	United States ( Texas)
Oxyacrum gracilis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Pagonia[73]	Gen. et sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Genus includes new species P. spiralis.
Palaeorhaphis? laevis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Paleofusus[73]	Gen. et 3 sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Genus includes new species P. bellus, P. parvicrassus and P. elegantissima.
Papuliscala? acus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Possibly a species of Papuliscala.
Papuliscala keani[110]	Sp. nov	Valid	Garvie	Eocene	Cook Mountain Formation	United States ( Texas)
Parasassia vargai[103]	Sp. nov	Valid	Vicián & Kovács	Eocene (Lutetian)	Csernye Formation	Hungary	A member of the family Cymatiidae.
Parthenina pacaudi[106]	Nom. nov	Valid	Landau & Micali	Miocene		France	A species of Parthenina; a replacement name for Pyrgulina (Parthenina) dollfusi Cossmann (1921).
Parvisipho? spiralis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Paskentana dixoni[87]	Sp. nov	Valid	Kaim et al.	Late Cretaceous (Cenomanian to Turonian)		Cyprus	A member of Abyssochrysoidea belonging to the family Paskentanidae.
Paskentana xenophontosi[87]	Sp. nov	Valid	Kaim et al.	Late Cretaceous (Cenomanian to Turonian)		Cyprus	A member of Abyssochrysoidea belonging to the family Paskentanidae.
Perotrochus bermotiensis[120]	Sp. nov	Valid	Bose, Das & Mondal	Oligocene		India	A species of Perotrochus.
Perrona harzhauseri[108]	Sp. nov	Valid	Kovács & Vicián	Middle Miocene	Lajta Limestone Formation	Hungary	A species of Perrona.
Perrona nemethi[108]	Sp. nov	Valid	Kovács & Vicián	Middle Miocene	Lajta Limestone Formation	Hungary	A species of Perrona.
Personopsis merlei[103]	Sp. nov	Valid	Vicián & Kovács	Eocene (Lutetian)	Csernye Formation	Hungary	A species of Personopsis.
Pfefferiola[126]	Nom. nov	Valid	Harzhauser & Neubauer	Eocene		Italy	A member of the family Streptaxidae; a replacement name for Oppenheimiella Pfeffer (1930).
Phos estotiensis[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Phos.
Pleurotomella irminonvilla[111]	Nom. nov	Valid	Pacaud	Eocene (Bartonian)		France	A species of Pleurotomella; a replacement name for Pleurotomella cossmanni Morellet & Morellet (1946).
Pliciscala obliqua[110]	Sp. nov	Valid	Garvie	Eocene	Cook Mountain Formation	United States ( Texas)
Poirieria (Paziella) delera[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Polinices tehuacana[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Polinices.
Popenoeum tabulatum[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Pressulasphaera[74]	Gen. et comb. nov	Valid	Karapunar & Nützel				A member of Pleurotomariida. The type species is "Pleurotomaria" pamphilus d'Orbigny (1850).
Priscoficus johnfoxi[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Priscoficus nodulina[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Proceraedis[73]	Gen. et 2 sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Genus includes new species P. imbricataria and P. tegumenis.
Prososthenia esuegirottii[137]	Sp. nov	Valid	Dominici & Forli	Pliocene (Zanclean)		Italy	A member of the family Hydrobiidae.
Protoplex? zsoldosi[103]	Sp. nov	Valid	Vicián & Kovács	Eocene (Lutetian)	Csernye Formation	Hungary	A member of the family Cymatiidae.
Proviviparus[138]	Gen. et sp. nov	Valid	Frese & Ponder	Late Jurassic	Talbragar Fish Bed	Australia	A member of the family Viviparidae. Genus includes new species P. talbragarensis
Pseudoananias[74]	Gen. et comb. nov	Valid	Karapunar & Nützel				A member of Pleurotomariida. The type species is "Pleurotomaria" subgranulata Münster (1841).
Pseudokatosira uhlhingleyorum[139]	Sp. nov	Valid	Gründel, Davies & Simonsen	Early Jurassic (Pliensbachian)
Pseudolatirus acuelatus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Pseudolatirus.
Pseudolatirus boettgeri[140]	Nom. nov	Valid	Kovács	Miocene		Romania	A member of the family Fasciolariidae; a replacement name for Fusus sublamellosus Boettger (1906). Originally described as a species of Pseudolatirus; Harzhauser, Landau & Vermeij (2024) tentatively placed it in the genus Takashius.[141]
Pseudosassia[103]	Gen. et comb. et 2 sp. nov	Valid	Vicián & Kovács	Paleocene–Miocene		Austria  Belgium  Bulgaria  Czech Republic  Denmark  France  Germany  Hungary  Italy  Netherlands  Poland  Romania  Serbia  Slovakia  Turkey  Ukraine  United Kingdom	A member of the family Cymatiidae. The type species is "Triton" flandricum Koninck (1838); genus also includes new species P. gurdoni and P. traceyi.
Pseudoterebellum[142]	Gen. et comb. nov	Valid	Maxwell, Rymer & Congdon	Eocene	Ghazij Shales	Indonesia  Pakistan  Saudi Arabia	A member of the family Seraphsidae. The type species is "Terebellum" diversiornatum Eames (1952); genus also includes "Terebellum" saudiensis Abbass (1972) and "Terebellum" olympiae Rolando (2001).
Ptychatractus cataractus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Ptychatractus.
Pulchraspira[113]	Gen. et sp. nov	In press	Yu, Salvador & Jarzembowski	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Genus includes new species P. teneristoma.
Puposyrnola? confusa[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Possibly a species of Puposyrnola.
Puposyrnola multibaca[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Puposyrnola.
Puposyrnola obliqua[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Puposyrnola.
Puposyrnola shirleyae[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Puposyrnola.
Pusia aturensis[91]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Pusia.
Pusia confunda[84]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Dej Formation	Romania	A species of Pusia.
Pusia? falsitranssylvanica[84]	Nom. nov	Valid	Harzhauser & Landau	Miocene	Dej Formation	Romania	Possibly a species of Pusia; a replacement name for Mitra (Costellaria) recticosta var. transsylvanica Boettger (1902).
Pusia subpyrenaica[91]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Pusia.
Pusia virodunensis[91]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Pusia.
Pustularia korneli[133]	Sp. nov	Valid	Fehse & Vicián	Miocene		Indonesia	A species of Pustularia.
Pyrgiscus elegans[143]	Sp. nov	Valid	Tabanelli et al.	Pliocene		Italy	A species of Pyrgiscus.
Pyrgiscus jaapmulderi[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Pyrgiscus.
Pyrgulina marliesae[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Pyrgulina.
Retusa (Cylichnina) praemoryi[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Retusa.
Rhaphaulus zhuoi[113]	Sp. nov	In press	Yu, Salvador & Jarzembowski	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar
Rinaldoella tornata[74]	Sp. nov	Valid	Karapunar & Nützel	Late Triassic	San Cassiano Formation	Italy	A member of Pleurotomariida.
Ringicula taenia[110]	Sp. nov	Valid	Garvie	Eocene	Cook Mountain Formation	United States ( Texas)	A species of Ringicula.
Sansonia texana[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Sassella[73]	Gen. et 2 sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Genus includes new species S. mixta and S. frosti.
Scalptia nemethi[144]	Sp. nov	Valid	Kovács & Vicián	Miocene	Lajta Formation	Hungary	A species of Scalptia.
Scaphella kendrae[145]	Sp. nov	Valid	Petuch & Berschauer	Pliocene (Zanclean)	Murdock Formation	United States ( Florida)	A species of Scaphella.
Schizogonium undae[74]	Sp. nov	Valid	Karapunar & Nützel	Late Triassic	San Cassiano Formation	Italy	A member of Pleurotomariida.
Seraphs kaindraperi[142]	Sp. nov	Valid	Maxwell, Rymer & Congdon	Eocene (Priabonian)		Jamaica	A member of the family Seraphsidae.
Similalvania[76]	Gen. et comb. nov	Valid	Chirli & Forli	Pliocene		Italy	A member of the family Rissoidae. The type species is "Alvania" magistra Chirli (2006)
Solariella microstriata[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Solariella.
Solariorbis planulatis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Squamoworthenia[127]	Gen. et comb. nov	Valid	Mazaev	Carboniferous		Australia  Germany  Kazakhstan	Genus includes S. duponti (Holzapfel, 1889) and "Worthenia" crenilunula Yoo (1994).
Stellaria striatissima[75]	Sp. nov	Valid	Raven	Miocene		Malaysia	A species of Stellaria.
Striacingulum[74]	Gen. et comb. nov	Valid	Karapunar & Nützel				A member of Pleurotomariida. The type species is "Pleurotomaria" cancellatocingulata Klipstein (1844).
Striaticostatum figus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Stuckenbergispira dombarensis[127]	Sp. nov	Valid	Mazaev	Early Carboniferous		Kazakhstan
Sveltella novus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Sveltella.
Sveltia sofiae[146]	Sp. nov	Valid	Landau & da Silva	Pliocene		Portugal	A species of Sveltia.
Syrnola debrae[110]	Sp. nov	Valid	Garvie	Eocene	Cook Mountain Formation	United States ( Texas)	A species of Syrnola.
Taioma? argilla[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Taioma calxa[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Taranis occulta[72]	Sp. nov	Valid	Brunetti & Cresti	Pliocene		Italy	A species of Taranis.
Teinostoma regularis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Teinostoma.
Terebellum humilispirum[75]	Sp. nov	Valid	Raven	Miocene		Malaysia	A species of Terebellum.
Terebra (Mirula) mexia[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Terebra.
Terebrifusus megahani[110]	Sp. nov	Valid	Garvie	Eocene	Cook Mountain Formation	United States ( Texas)
Thylacodes devriesi[147]	Sp. nov	In press	Sanfilippo et al.	Miocene		Peru	A species of Thylacodes.
Tornatellaea globosa[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Tornatellaea linifera[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Tripsycha strapplei[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Tritia miniridibunda[86]	Sp. nov	Valid	Lozouet	Miocene		France	A species of Tritia.
Tritia praebasteroti[86]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Tritia.
Tritiaria formosa[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Trivia disjuncta[136]	Sp. nov	Valid	Fehse	Late Oligocene		France	A species of Trivia.
Trivia simplex[136]	Sp. nov	Valid	Fehse	Late Oligocene		France	A species of Trivia.
Trona colossus[118]	Sp. nov	Valid	Groves & Landau	Early–middle Miocene	Baitoa Formation	Dominican Republic	A species of Trona.
Truncatella jiaozhouensis[148]	Sp. nov	In press	Yu et al.	Late Cretaceous	Jiaolai Basin	China	A species of Truncatella.
Tumidosipho[73]	Gen. et sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	Genus includes new species T. tehuacana.
Turbo (Marmarostoma) ishidai[149]	Sp. nov	Valid	Tomida, Sano & Kase	Miocene	Ena Limestone	Japan	A species of Turbo.
Turbonilla bincki[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Turbonilla.
Turbonilla bongiardinoi[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Turbonilla.
Turbonilla crovatoi[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Turbonilla.
Turbonilla malacitana[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Turbonilla.
Turbonilla mauroi[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Turbonilla.
Turbonilla plioalboranensis[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Turbonilla.
Turbonilla velerinensis[106]	Sp. nov	Valid	Landau & Micali	Pliocene		Spain	A species of Turbonilla.
Turricula frosti[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Turricula.
Turricula imbricata[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Turricula.
"Turritella" calax[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Umbilia furneauxensis[150]	Sp. nov	Valid	Southgate, Militz & Roberts	Pliocene		Australia	A species of Umbilia.
Unitas duomillias[110]	Sp. nov	Valid	Garvie	Eocene	Cook Mountain Formation	United States ( Texas)
Unitas euani[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Unitas petiti[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Valvata jiaolaiensis[148]	Sp. nov	In press	Yu et al.	Late Cretaceous	Jiaolai Basin	China	A species of Valvata.
Vanikoropsis bashforthi[151]	Sp. nov	Valid	Schnetler & Nielsen	Paleocene (Danian)	Agatdal Formation	Greenland	Possibly a member of the family Vanikoridae.
Vanikoropsis jakobseni[151]	Sp. nov	Valid	Schnetler & Nielsen	Paleocene (Danian)	Agatdal Formation	Greenland	Possibly a member of the family Vanikoridae.
Vanikoropsis mortenseni[151]	Sp. nov	Valid	Schnetler & Nielsen	Paleocene (Selandian)	Kerteminde Marl	Denmark	Possibly a member of the family Vanikoridae.
Varicospira bekenuensis[75]	Sp. nov	Valid	Raven	Miocene		Malaysia	A species of Varicospira.
Varicospira reticulata[75]	Sp. nov	Valid	Raven	Miocene		Malaysia	A species of Varicospira.
Vermicularia? columella[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Vexillum transalpinum[84]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Baden Formation	Austria  Hungary  Romania  Turkey	A species of Vexillum.
Vitrinella discus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Vitrinella.
Vitrinella praelaevis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Vitrinella.
Volutifusus dougsheltoni[145]	Sp. nov	Valid	Petuch & Berschauer	Pliocene (Piacenzian)	Tamiami Formation	United States ( Florida)	A species of Volutifusus.
Volutifusus kissimmeensis[145]	Sp. nov	Valid	Petuch & Berschauer	Pliocene (Piacenzian)	Tamiami Formation	United States ( Florida)	A species of Volutifusus.
Volutomorpha ultima[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Wormsina[102]	Gen. et comb. nov	Valid	Harzhauser & Landau	Miocene		Romania	A member of the family Mitridae. The type species is "Mitra (Cylindra)" transsylvanica Hoernes & Auinger (1880).
Wortheniella klipsteini[74]	Sp. nov	Valid	Karapunar & Nützel	Late Triassic	San Cassiano Formation	Italy	A member of Pleurotomariida.
Wortheniella paolofedelei[74]	Sp. nov	Valid	Karapunar & Nützel	Late Triassic	San Cassiano Formation	Italy	A member of Pleurotomariida.
Zafrona sphaerocorrugata[89]	Sp. nov	Valid	Harzhauser & Landau	Miocene	Baden Formation	Austria  Bosnia and Herzegovina  Bulgaria  Hungary  Romania  Ukraine	A species of Zafrona.
Zebinella tehuacana[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Zebinella.
Zonaria vokesae[118]	Sp. nov	Valid	Groves & Landau	Early–middle Miocene	Baitoa Formation	Dominican Republic	A species of Zonaria.

Bivalves

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Acar beadata[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Acar.
Amphidonte mirkamalovi[152]	Sp. nov	Valid	Metelkin & Kosenko	Early Cretaceous (Aptian)			An oyster.
Arcopsis subrhombus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Arcopsis tumulus[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Arkhanguelskya[153]	Gen. et comb. nov	Valid	Cooper & Leanza	Cretaceous			A member of the family Buchotrigoniidae. Genus includes "Trigonia" weberi Arkhangelsky (1916).
Baluchicardia? eoplata[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Bengtsonigonia[154]	Nom. nov	Valid	Ceccolini & Cianferoni	Early Cretaceous	Riachuelo Formation	Brazil	A replacement name for Bengtsonella Cooper & Leanza (2019).
Bohemigonia[155]	Gen. et sp. et comb. nov	Valid	Cooper			Czech Republic	A member of the family Megatrigoniidae. The type species is B. frici; genus also includes "Trigonia" debuchii Geinitz (1840).
Caryocorbula? praefigera[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Chama ferrata[156]	Sp. nov	Valid	Berezovsky	Eocene		Ukraine	A species of Chama.
Chama subcalcarata[156]	Sp. nov	Valid	Berezovsky	Eocene		Ukraine	A species of Chama.
Corbulamella palaeocenica[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Crassatellina minuta[157]	Sp. nov	Valid	Jaitly et al.	Late Cretaceous (Turonian)	Nodular Limestone Formation	India	A member of the family Lucinidae belonging to the subfamily Eriphylinae.
Crassatellina paleocaenica[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Cuspidaria texana[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Cuspidaria.
Diplodonta frosti[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Diplodonta paleocaenica[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Dorbigonia[155]	Gen. et comb. nov	Valid	Cooper	Cretaceous		France	A member of the family Megatrigoniidae. Genus includes "Trigonia" disparilis d'Orbigny (1844).
Elianegonia[154]	Nom. nov	Valid	Ceccolini & Cianferoni	Late Cretaceous		Madagascar	A replacement name for Elianella Cooper (2015).
Eomiltha strangensis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Eurydesma sacculum snapperensis[158]	Ssp. nov	Valid	Waterhouse	Permian	Wandrawandian Formation	Australia	A bivalve belonging to the group Pectinida and the family Eurydesmidae.
Freneixicardia picturata[159]	Sp. nov	Valid	Berezovsky	Eocene		Ukraine	A cockle.
Frenguelliella eopacifica[160]	Sp. nov	Valid	Echevarría, Damborenea & Manceñido	Early Jurassic		Argentina	A member of Trigoniida belonging to the family Frenguelliellidae.
Glycymeris laevigata[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Glycymeris.
Grammatodon superstes[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Gregariella frosti[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Gregariella.
Himenourella[155]	Gen. et comb. nov	Valid	Cooper	Cretaceous		Japan  Russia	A member of the family Megatrigoniidae. Genus includes "Trigonia" jimbonis Strand (1928), "Trigonia" sawatai Yehara (1923), "Apiotrigonia" subjimboi Miroljubov in Vereshchagin et al. (1965) and "Trigonia" subovalis Jimbo (1894).
Kolymia rutskovi[161]	Sp. nov	Valid	Biakov in Biakov et al.	Permian		Russia
Kushmurunskya[155]	Gen. et comb. nov	Valid	Cooper	Cretaceous		Kazakhstan	A member of the family Megatrigoniidae. Genus includes "Trigonia" gladkowskii Shcheglova-Borodina (1960).
Kyzylkumgonia[155]	Gen. et comb. nov	Valid	Cooper	Cretaceous			A member of the family Megatrigoniidae. Genus includes "Trigonia" syrdariensis Arkhangelsky (1916), "Trigonia" turcmenensis Arkhangelsky (1916) and "Megatrigonia" tagamensis Belyakova (1980).
Loxocardium marmoreum[159]	Sp. nov	Valid	Berezovsky	Eocene		Ukraine	A cockle.
Lucina crassiplica[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Lucina.
Lucina multivittae[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Lucina.
Lucina tehuacanensis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Lucina.
Maedagonia[155]	Gen. et comb. et nom. nov	Valid	Cooper	Cretaceous		Japan  Russia	A member of the family Megatrigoniidae. Genus includes "Apiotrigonia" mikasaensis Tashiro (1979), "Apiotrigonia" ashizawaensis Saito (1962), "Apiotrigonia" crassoradiata Nakano (1957), "Apiotrigonia" futabaensis Maeda & Kawabe (1967), "Apiotrigonia" hetonaiana Tashiro (1978), "Apiotrigonia" hironoensis Maeda & Kawabe (1967), "Apiotrigonia" jimboi Nakano (1957), "Apiotrigonia" obliquecostata Nakano (1957), "Apiotrigonia" obsoleta Nakano (1957), "Apiotrigonia" orikiensis Maeda & Kawabe (1967), "Apiotrigonia" undulosa Nakano (1957) and "Apiotrigonia" utoensis Tashiro (1972), as well as M. yabei (a replacement name for Trigonia subovalis var. minor Yabe & Nagao, 1925).
Martesia mazanticus[162]	Sp. nov	Valid	Castañeda-Posadas, Trujillo-Hernández & Zúñiga-Mijangos	Miocene	Mexican amber	Mexico	A species of Martesia.
Matsudagonia[155]	Gen. et comb. nov	Valid	Cooper	Cretaceous		Japan	A member of the family Megatrigoniidae. Genus includes "Apiotrigonia" dubia Tashiro (1979).
Moerickella[160]	Gen. nov	Valid	Echevarría, Damborenea & Manceñido	Early Jurassic		Argentina	A member of Trigoniida, most likely the oldest member of the family Myophorellidae.
Naiadites devonicus[163]	Sp. nov	Valid	Scholze & Gess	Devonian (Famennian)	Witpoort Formation	South Africa
Opis reticulata[157]	Sp. nov	Valid	Jaitly et al.	Late Cretaceous (Turonian)	Nodular Limestone Formation	India	A member of the family Lucinidae belonging to the subfamily Opinae.
Ostrea jibananandai[164]	Sp. nov	Valid	Halder & Mitra	Eocene (Ypresian)	Cambay Shale	India	A species of Ostrea.
Palaeolignopholas[165]	Gen. et sp. nov	Valid	Bolotov et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Pholadidae belonging to the subfamily Martesiinae. The type species is P. kachinensis.
Pandora? paleocenica[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Parmicorbula? sublaevis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Perezigonia[154]	Nom. nov	Valid	Ceccolini & Cianferoni	Middle Jurassic		Chile	A replacement name for Perezella Cooper (2015).
Pojetaia robsonae[166]	Sp. nov	Valid	Peel & Skovsted	Cambrian Stage 4	Kane Basin Formation	Canada ( Nunavut)
Porterius? strangensis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Poultoniella[155]	Gen. et comb. nov	Valid	Cooper	Cretaceous		Canada ( British Columbia)	A member of the family Megatrigoniidae. Genus includes "Trigonia" newcombei Packard (1921).
Promyophorella basoaltorum[160]	Sp. nov	Valid	Echevarría, Damborenea & Manceñido	Early Jurassic		Argentina	A member of Trigoniida belonging to the family Myophorellidae.
Promyophorella? sanjuanina[160]	Sp. nov	Valid	Echevarría, Damborenea & Manceñido	Early Jurassic		Argentina	A member of Trigoniida belonging to the family Myophorellidae.
Pseudapiotrigonia[155]	Gen. et comb. nov	Valid	Cooper			United States ( South Dakota  Utah)	A member of the family Vaugoniidae. Genus includes "Trigonia" sturgisensis Whitfield & Hovey (1906), as well as "Vaugonia" utahensis Imlay (1964).
Pseudovaugonia[160]	Gen. nov	Valid	Echevarría, Damborenea & Manceñido	Early Jurassic		Argentina	A member of Trigoniida belonging to the family Myophorellidae.
Pteromeris nodulina[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Pteromeris.
Repmanigonia[154]	Nom. nov	Valid	Ceccolini & Cianferoni	Jurassic		Russia	A replacement name for Repmania Kolesnikov (1980).
Schedocardia imperfecta[159]	Sp. nov	Valid	Berezovsky	Eocene		Ukraine	A cockle.
Sinoliaoningia[154]	Nom. nov	Valid	Ceccolini & Cianferoni	Late Triassic	Yangcaogou Formation	China	A replacement name for Liaoningia Yu & Dong (1993).
Sokolovskya[155]	Gen. et comb. nov	Valid	Cooper				A member of the family Megatrigoniidae. Genus includes "Apiotrigonia" levinae Pojarkova (1990) and "Apiotrigonia" khoresmensis Belyakova (1965).
Sonoragonia[155]	Gen. et comb. et nom. nov	Valid	Cooper	Early Cretaceous	Lowell Formation	United States ( Arizona)	A member of the family Megatrigoniidae. Genus includes "Trigonia" cragini Stoyanow (1949), as well as S. scotti (a replacement name for Trigonia kitchini Stoyanow, 1949).
Sphenotomorpha orientalis[167]	Sp. nov	Valid	Dalenz Farjat	Devonian (Pragian)	Icla Formation	Bolivia	A member of Modiomorphida belonging to the family Modiomorphidae.
Sunetta crassatelliformis[168]	Sp. nov	Valid	Haga & Fukuda in Fukuda et al.	Early Pleistocene		Japan	A species of Sunetta.
Sunetta nomurai[168]	Sp. nov	Valid	Haga & Fukuda in Fukuda et al.	Pleistocene		Taiwan	A species of Sunetta.
Syndesmyella pliominor[169]	Sp. nov	Valid	Brunetti & Cresti	Early Pliocene		Spain	A member of the family Semelidae.
Tellina brevis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A species of Tellina.
Terrypoultoniella[160][170]	Gen. et sp. nov	Valid	Echevarría, Damborenea & Manceñido	Early Jurassic (Pliensbachian–Toarcian)		Argentina  Portugal?	A member of Trigoniida belonging to the family Frenguelliellidae. The type species is "Poultoniella" jaworskii Echevarría, Damborenea & Manceñido (2021); genus might also include "Trigonia" malladae Choffat (1885).
Tindaria hamuroi[171]	Sp. nov	Valid	Amano & Haga	Miocene	Higashibessho Formation	Japan	A member of the family Tindariidae.
Titomaya[172]	Gen. et comb. et sp. nov	Valid	Alvarez & del Río	Paleocene (Danian)	Lefipán Formation Salamanca Formation	Argentina	A member of Venerida belonging to the group Veneroidea. Genus includes "Meretrix" chalcedonica, as well as new species T. longobucca.
Tivelina texana[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Tivelina trigonalis[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)
Unionites kobyumensis[173]	Sp. nov	Valid	Biakov in Biakov & Kutygin	Late Permian		Russia
Vaccinites alencasteri[174]	Sp. nov	Valid	Mitchell	Late Cretaceous (probably late Turonian or early Coniacian)	Cuautla Formation	Mexico	A rudist bivalve.
Vaccinites temazcali[174]	Sp. nov	Valid	Mitchell	Late Cretaceous (Campanian)	Temazcal Limestone	Mexico	A rudist bivalve.
Villamilia[153]	Gen. et comb. nov	Valid	Cooper & Leanza	Cretaceous		Colombia	A member of the family Buchotrigoniidae. Genus includes "Buchotrigonia" etayoi Villamil (1992), "Trigonia" humboldtii Buch (1839), "Buchotrigonia" kauffmani Villamil (1992), "Buchotrigonia" perezi Villamil (1992) and "Vaugonia niranohamensis" santamariae Geyer (1973; raised to the rank of the species V. santamariae).
Weyla santuccii[175]	Sp. nov	Valid	Damborenea et al.	Early Jurassic (Hettangian)	Pogibshi Formation	United States ( Alaska)	A scallop.
Whitfieldiella luceae[174]	Sp. nov	Valid	Mitchell	Late Cretaceous (Campanian)	Clifton Limestone Cotui Limestone	Jamaica  United States ( Puerto Rico)	A rudist bivalve.
Yunannia[176]	Gen. et 2 sp. nov	Valid	Zhang & Niu in Zhang et al.	Ordovician	Dongchong Formation	China	A member of Pteriomorphia of uncertain phylogenetic placement. The type species is Y. gankengensis; genus also includes Y. yunkaiensis.

Research

• A study on the evolution and possible causes of the extinction of alatoconchid bivalves is published by Chen et al. (2021).[177]
• A study on the arcoid bivalve diversity during the Eocene doubthouse interval of global climate cooling is published by Hickman (2021).[178]

Other molluscs

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Anabarella? navaranae[179]	Sp. nov	Valid	Peel	Cambrian Stage 4	Aftenstjernesø Formation	Greenland
Cadulus praepalmerae[73]	Sp. nov	Valid	Garvie	Paleocene (Danian)	Kincaid Formation	United States ( Texas)	A tusk shell, a species of Cadulus.
Catalanispira prima[105]	Sp. nov	Valid	Ebbestad & Cope	Ordovician (Floian)		United Kingdom	A member of the family Onychochilidae.
Figurina? polaris[179]	Sp. nov	Valid	Peel	Cambrian Stage 4	Aftenstjernesø Formation	Greenland	A helcionellid. Subsequently made the type species of a separate genus Sermeqiconus.[180]
Hippopharangites groenlandicus[179]	Sp. nov	Valid	Peel	Cambrian Stage 4	Aftenstjernesø Formation	Greenland	A member of the stem group of Aculifera.
Holoornatus[181]	Gen. et sp. nov	In press	Zhang et al.	Cambrian (Fortunian)		China	A maikhanellid. Genus includes new species H. strigatus.
Inughuitoconus[179]	Gen. et sp. nov	Valid	Peel	Cambrian Stage 4	Aftenstjernesø Formation	Greenland	A member of the stem group of Aculifera. Genus includes new species I. borealis.
Lepidozona luzanovkensis[182]	Sp. nov	Valid	Sirenko	Paleocene	Luzanovka Beds	Ukraine	A chiton, a species of Lepidozona.
Minodentalium[183]	Gen. et 2 sp. et comb. nov	Valid	Asato & Kase	Permian	Akasaka Limestone	Belgium  Japan  New Zealand  United States	A tusk shell. Genus includes new species M. hayasakai and M. okumurai, as well as "Plagioglypta" furcata Waterhouse (1980), "Pl." girtyi Knight (1940), "Pl." subannulata Easton (1962), "Dentalium" ingens De Koninck (1843), "D." meekianum Geinitz (1866), "Pl." prosseri Morningstar (1922), "Dentalium" priscum Münster in Goldfuss (1842) and "D." herculeum De Koninck (1863).
Ocruranus? kangerluk[179]	Sp. nov	Valid	Peel	Cambrian Stage 4	Aftenstjernesø Formation	Greenland	A member of the stem group of Aculifera.
Prodentalium onoi[183]	Sp. nov	Valid	Asato & Kase	Permian	Akasaka Limestone	Japan	A tusk shell.
Progadilinoides[184]	Gen. et sp. nov	Valid	Guzhov	Early Cretaceous (Aptian)		Russia	A tusk shell belonging to the family Pulsellidae. The type species is P. balakovensis.
Pseudomyona groenlandica[185]	Sp. nov	Valid	Peel	Cambrian (Drumian)	Fimbuldal Formation	Greenland	A member of Rostroconchia belonging to the group Tuarangiida and the family Tuarangiidae.
Qingjianglepas[186]	Gen. et sp. nov	In press	Li et al.	Cambrian		China	A minute limpet-shaped shell, probably belonging to a helcionelloid. Genus includes new species Q. elegans.
Stenotheca? higginsi[179]	Sp. nov	Valid	Peel	Cambrian Stage 4	Aftenstjernesø Formation	Greenland

General research

• A study on the relationship between temperature and diversity in the fossil record of marine molluscs over the last 145 million years is published by Boag, Gearty & Stockey (2021).[187]
• A study on the relationship between the species richness of shallow-marine molluscs, the number of ecological functional groups, and oceanic temperature in the fossil record of New Zealand over the last ~40 million years is published by Womack et al. (2021).[188]

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