^Penz, C. M., Devries, P. J., & Wahlberg, N. 2012. Diversification of Morpho butterflies (Lepidoptera, Nymphalidae): a re‐evaluation of morphological characters and new insight from DNA sequence data. Systematic Entomology, 37(4), 670-685. (英文)
^Penz CM, and DeVries PJ. 2002. Phylogenetic analysis of Morpho butterfles (Nymphalidae, Morphinae): implications for classification and natural history. Amer. Mus. Novitates 3374: 1-33.
參考文獻
Chazot, N., Panara, S., Zilbermann, N., Blandin, P., Le Poul, Y., Cornette, R., ... & Debat, V. 2016. Morpho morphometrics: Shared ancestry and selection drive the evolution of wing size and shape in Morpho butterflies. Evolution. (英文)
Blandin, P., & Purser, B. 2013. Evolution and diversification of Neotropical butterflies: insights from the biogeography and phylogeny of the genus Morpho Fabricius, 1807 (Nymphalidae: Morphinae), with a review of the geodynamics of South America. Tropical Lepidoptera Research, 23(2), 62-85. (英文)
Cassildé, C., P. Blandin & J.F. Silvain. 2012. Phylogeny of the genus Morpho Fabricius 1807: insights from two mitochondrial genes (Lepidoptera: Nymphalidae). Ann. Soc. entomol. Fr. (n.s.), 48: 173-188. (英文)
Cassildé, C., Blandin, P. Pierre, J. & T. Bourgoin. 2010. Phylogeny of the genus Morpho Fabricius, 1807, revisited (Lepidoptera, Nymphalidae). Bull. Soc. ent. Fr., 115: 225-250. (英文)
Le Moult E, and Real P. 1962. Les Morpho d'Amérique du Sud et Centrale. Editions scientifiques du Cabinet entomologique E. Le Moult, Paris.
寿建新 周尧 李宇飞. 世界蝴蝶分類名錄. 中國: 陝西埔學技術出版社. 2006-04-01. ISBN 9787536936768(中文(简体)).