^Blanca Huertas, Gerardo Lamas, Giovanny Fagua, James Mallet, Shinichi Nakahara and Keith Willmott. 2016. A Remarkable New Butterfly Species from western Amazonia (Lepidoptera: Nymphalidae: Satyrinae). [Una notable nueva especie de mariposa de la Amazonía occidental (Lepidoptera: Nymphalidae: Satyrinae)]. Conservación Colombiana. No. 24 – 27; 5-11.
參考文獻
Forster W. 1964. Beiträge zur Kenntnis der Insektenfauna Boliviens. XIX. Einleitung Lepidoptera III. Satyridae. Veröffentlichungen der zoologischen Staatsammlung München 3: 81-160, plates 25-33.
Lamas G ed. 2004. Atlas of Neotropical Lepidoptera. Checklist: Part 4A Hesperioidea - Papiionoidea. Gainesville: Scientific Publishers/Association of Tropical Lepidoptera.
Miller LD. 1968. The higher classification, phylogeny and zoogeography of the Satyridae (Lepidoptera). Mem. Amer. Entomol. Soc. 24: 1-174.
Murray D, and Prowell DP. 2005. Molecular phylogenetics and evolutionary history of the neotropical satyrine subtribe Euptychiina (Nymphalidae: Satyrinae). Mol. Phylogenet. Evol. 34: 67-80.
Peña C, Nylin S, Freitas AVL and Wahlberg N. 2010. Biogeographical history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae). Zoologica Scripta 39: 243-258.
Peña C, Wahlberg N, Weingartner E, Kodandaramaiah U, Nylin S, Freitas AVL, and Brower AVZ. 2006. Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Mol. Phylogenet. Evol. 40: 29-49.
近期研究
ZACCA, T., CASAGRANDE, M. M., MIELKE, O. H., HUERTAS, B., ESPELAND, M., FREITAS, A. V., ... & LAMAS, G. (2020). Revalidation of Vareuptychia Forster, 1964, description of Vanima gen. nov., and notes on Euptychia cleophes Godman & Salvin, 1889 (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa, 4858(1), 1-34.
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