Peña, C. & Wahlberg, N. 2008 Prehistorical climate change increased diversification of a group of butterflies. Biology Letters (Royal Society) 4, 274-278.
Corbet AS, Pendlebury HM, and Eliot JN. 1992. The butterflies of the Malay Peninsula. Malayan Nature Society, Kuala Lumpur.
Parsons M. 1999. The butterflies of Papua New Guinea: their systematics and biology. Academic Press, San Diego.
Pena C, Wahlberg N, Weingartner E, Kodandaramaiah U, Nylin S, Freitas AVL, and Brower AVZ. 2006. Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Mol. Phylogenet. Evol. 40: 29-49.
寿建新 周尧 李宇飞. 世界蝴蝶分類名錄. 中國: 陝西科學技術出版社. 2006-04-01. ISBN 9787536936768(中文(简体)).