^Ksepka, Daniel T.; Field, Daniel J.; Heath, Tracy A.; Pett, Walker; Thomas, Daniel B.; Giovanardi, Simone; Tennyson, Alan J. D. Largest-known fossil penguin provides insight into the early evolution of sphenisciform body size and flipper anatomy. Journal of Paleontology. 8 February 2023: 1–20. ISSN 0022-3360. S2CID 256709376. doi:10.1017/jpa.2022.88(英语).
^Daniel T. Ksepka, Daniel J. Field, Tracy A. Heath, Walker Pett, Daniel B. Thomas, Simone Giovanardi and Alan J.D. Tennyson. 2023. Largest-known Fossil Penguin provides insight into the early Evolution of sphenisciform Body Size and Flipper Anatomy. Journal of Paleontology. First View , pp. 1 - 20
DOI: 10.1017/jpa.2022.88
UUID: zoobank.org/15b1d5b2-a5a0-4aa5-ba0a-8ef3b8461730