Prosody (linguistics)

In linguistics, prosody (/ˈprɒsədi, ˈprɒz-/)[1][2] is the study of elements of speech that are not individual phonetic segments (vowels and consonants) but which are properties of syllables and larger units of speech, including linguistic functions such as intonation, stress, and rhythm. Such elements are known as suprasegmentals.[3]

Prosody reflects the nuanced emotional features of the speaker or of their utterances: their obvious or underlying emotional state, the form of utterance (statement, question, or command), the presence of irony or sarcasm, certain emphasis on words or morphemes, contrast, focus, and so on. Prosody displays elements of language that are not encoded by grammar, punctuation or choice of vocabulary.

Attributes of prosody

In the study of prosodic aspects of speech, it is usual to distinguish between auditory measures (subjective impressions produced in the mind of the listener) and objective measures (physical properties of the sound wave and physiological characteristics of articulation that may be measured objectively). Auditory (subjective) and objective (acoustic and articulatory) measures of prosody do not correspond in a linear way.[4] Most studies of prosody have been based on auditory analysis using auditory scales.

Auditorily, the major prosodic variables are:

  • pitch of the voice (varying between low and high)
  • length of sounds (varying between short and long)
  • loudness, or prominence (varying between soft and loud)
  • timbre or phonatory quality (quality of sound)

Acoustically, these prosodic variables correspond closely to:

Visualization of the prosody of a male voice saying "speech prosody": pitch in ribbon height, and periodic energy in ribbon width and darkness.
Audio for the visualization above.
  • fundamental frequency (measured in hertz, or cycles per second)
  • duration (measured in time units such as milliseconds or seconds)
  • intensity, or sound pressure level (measured in decibels)
  • spectral characteristics (distribution of energy at different parts of the audible frequency range)

Different combinations of these variables are exploited in the linguistic functions of intonation and stress, as well as other prosodic features such as rhythm and tempo.[4] Additional prosodic variables have been studied, including voice quality and pausing. The behavior of the prosodic variables can be studied either as contours across the prosodic unit or by the behavior of boundaries.[5]

Phonology

Prosodic features are suprasegmental, since they are properties of units of speech that are defined over groups of sounds rather than single segments.[6] When talking about prosodic features, it is important to distinguish between the personal characteristics that belong to an individual's voice (for example, their habitual pitch range, intonation patterns, etc.) and the independently variable prosodic features that are used contrastively to communicate meaning (for example, the use of changes in pitch to indicate the difference between statements and questions). Personal characteristics that belong to an individual are not linguistically significant while prosodic features are. Prosody has been found across all languages and is described to be a natural component of language. The defining features of prosody that display the nuanced emotions of an individual differ across languages and cultures.

Intonation

Some writers (e.g., O'Connor and Arnold)[7] have described intonation entirely in terms of pitch, while others (e.g., Crystal) [8] propose that "intonation" is a combination of several prosodic variables. English intonation is often said to be based on three aspects:

  • The division of speech into units
  • The highlighting of particular words and syllables
  • The choice of pitch movement (e.g., fall or rise)

The choice of pitch movement and the highlighting of particular words to create different intonation patterns can be seen in the following English conversation:

"That's a cat?"
"Yup. That's a cat."
"A cat? I thought it was a mountain lion!"

The exchange above is an example of using intonation to highlight particular words and to employ rising and falling of pitch to change meaning. If read out loud, the pitch of the voice moves in different directions on the word "cat." In the first line, pitch goes up, indicating a question. In the second line, pitch falls, indicating a statement— a confirmation of the first line in this case. Finally, in the third line, a complicated rise-fall pattern indicates incredulity. Each pitch/intonation pattern communicates a different meaning.[6]

An additional pitch-related variation is pitch range; speakers are capable of speaking with a wide range of pitch (this is usually associated with excitement), while at other times with a narrow range. English makes use of changes in key; shifting one's intonation into the higher or lower part of one's pitch range is believed to be meaningful in certain contexts.[9][10]

Stress

Stress functions as the means of making a syllable prominent. Stress may be studied in relation to individual words (named "word stress" or lexical stress) or in relation to larger units of speech (traditionally referred to as "sentence stress" but more appropriately named "prosodic stress"). Stressed syllables are made prominent by several variables. Stress is typically associated with the following:

  • pitch prominence (a pitch level that is different from that of neighboring syllables, or a pitch movement)
  • increased length (duration)
  • increased loudness (dynamics)
  • differences in timbre: in English and some other languages, stress is associated with aspects of vowel quality (whose acoustic correlate is the formant frequencies or spectrum of the vowel). Unstressed vowels tend to be centralized relative to stressed vowels, which are normally more peripheral in quality[11]

Some of these cues are more powerful or prominent than others. Alan Cruttenden, for example, writes "Perceptual experiments have clearly shown that, in English at any rate, the three features (pitch, length and loudness) form a scale of importance in bringing syllables into prominence, pitch being the most efficacious, and loudness the least so".[12]

When pitch prominence is the major factor, the resulting prominence is often called accent rather than stress.[13]

There is considerable variation from language to language concerning the role of stress in identifying words or in interpreting grammar and syntax.[14]

Tempo

Rhythm

Although rhythm is not a prosodic variable in the way that pitch or loudness are, it is usual to treat a language's characteristic rhythm as a part of its prosodic phonology. It has often been asserted that languages exhibit regularity in the timing of successive units of speech, a regularity referred to as isochrony, and that every language may be assigned one of three rhythmical types: stress-timed (where the durations of the intervals between stressed syllables is relatively constant), syllable-timed (where the durations of successive syllables are relatively constant) and mora-timed (where the durations of successive morae are relatively constant). As explained in the isochrony article, this claim has not been supported by scientific evidence.

Pause

Voiced or unvoiced, the pause is a form of interruption to articulatory continuity such as an open or terminal juncture. Conversation analysis commonly notes pause length. Distinguishing auditory hesitation from silent pauses is one challenge. Contrasting junctures within and without word chunks can aid in identifying pauses.

There are a variety of "filled" pause types. Formulaic language pause fillers include "Like", "Er" and "Um", and paralinguistic expressive respiratory pauses include the sigh and gasp.

Although related to breathing, pauses may contain contrastive linguistic content, as in the periods between individual words in English advertising voice-over copy sometimes placed to denote high information content, e.g. "Quality. Service. Value".

Chunking

Pausing or its lack contributes to the perception of word groups, or chunks. Examples include the phrase, phraseme, constituent or interjection. Chunks commonly highlight lexical items or fixed expression idioms. Chunking prosody[15] is present on any complete utterance and may correspond to a syntactic category, but not necessarily. The well-known English chunk "Know what I mean?" in common usage sounds like a single word ("No-wada-MEEN?") due to blurring or rushing the articulation of adjacent word syllables, thereby changing the potential open junctures between words into closed junctures.

Functions

Prosody has had a number of perceptually significant functions in English and other languages, contributing to the recognition and comprehension of speech.[16]

Grammar

It is believed that prosody assists listeners in parsing continuous speech and in the recognition of words, providing cues to syntactic structure, grammatical boundaries and sentence type. Boundaries between intonation units are often associated with grammatical or syntactic boundaries; these are marked by such prosodic features as pauses and slowing of tempo, as well as "pitch reset" where the speaker's pitch level returns to the level typical of the onset of a new intonation unit. In this way potential ambiguities may be resolved. For example, the sentence "They invited Bob and Bill and Al got rejected" is ambiguous when written, although addition of a written comma after either "Bob" or "Bill" will remove the sentence's ambiguity. But when the sentence is read aloud, prosodic cues like pauses (dividing the sentence into chunks) and changes in intonation will reduce or remove the ambiguity.[17] Moving the intonational boundary in cases such as the above example will tend to change the interpretation of the sentence. This result has been found in studies performed in both English and Bulgarian.[18] Research in English word recognition has demonstrated an important role for prosody.[19][20]

Focus

Intonation and stress work together to highlight important words or syllables for contrast and focus.[21] This is sometimes referred to as the accentual function of prosody. A well-known example is the ambiguous sentence "I never said she stole my money", where there are seven meaning changes depending on which of the seven words is vocally highlighted.[22]

Discourse and pragmatic functions

Prosody helps convey many other pragmatic functions, including expressing attitudes (approval, uncertainty, dissatisfaction, and so on), flagging turn-taking intentions (to hold the floor, to yield the turn, to invite a backchannel like uh-huh, and so on), and marking topic structure (starting a new topic, closing a topic, interpolating a parenthetical remark, and so on), among others.[23][24][25] For example, David Brazil and his associates studied how intonation can indicate whether information is new or already established; whether a speaker is dominant or not in a conversation; and when a speaker is inviting the listener to make a contribution to the conversation.[26]

Emotion

Prosody is also important in signalling emotions and attitudes. When this is involuntary (as when the voice is affected by anxiety or fear), the prosodic information is not linguistically significant. However, when the speaker varies their speech intentionally, for example to indicate sarcasm, this usually involves the use of prosodic features. The most useful prosodic feature in detecting sarcasm is a reduction in the mean fundamental frequency relative to other speech for humor, neutrality, or sincerity. While prosodic cues are important in indicating sarcasm, context clues and shared knowledge are also important.[27]

Emotional prosody was considered by Charles Darwin in The Descent of Man to predate the evolution of human language: "Even monkeys express strong feelings in different tones – anger and impatience by low, – fear and pain by high notes."[28] Native speakers listening to actors reading emotionally neutral text while projecting emotions correctly recognized happiness 62% of the time, anger 95%, surprise 91%, sadness 81%, and neutral tone 76%. When a database of this speech was processed by computer, segmental features allowed better than 90% recognition of happiness and anger, while suprasegmental prosodic features allowed only 44%–49% recognition. The reverse was true for surprise, which was recognized only 69% of the time by segmental features and 96% of the time by suprasegmental prosody.[29] In typical conversation (no actor voice involved), the recognition of emotion may be quite low, of the order of 50%, hampering the complex interrelationship function of speech advocated by some authors.[30] However, even if emotional expression through prosody cannot always be consciously recognized, tone of voice may continue to have subconscious effects in conversation. This sort of expression stems not from linguistic or semantic effects, and can thus be isolated from traditional[clarification needed] linguistic content.[dubiousdiscuss] Aptitude of the average person to decode conversational implicature of emotional prosody has been found to be slightly less accurate than traditional facial expression discrimination ability; however, specific ability to decode varies by emotion. These emotional[clarification needed] have been determined to be ubiquitous across cultures, as they are utilized and understood across cultures. Various emotions, and their general experimental identification rates, are as follows:[31]

  • Anger and sadness: High rate of accurate identification
  • Fear and happiness: Medium rate of accurate identification
  • Disgust: Poor rate of accurate identification

The prosody of an utterance is used by listeners to guide decisions about the emotional affect of the situation. Whether a person decodes the prosody as positive, negative, or neutral plays a role in the way a person decodes a facial expression accompanying an utterance. As the facial expression becomes closer to neutral, the prosodic interpretation influences the interpretation of the facial expression. A study by Marc D. Pell revealed that 600 ms of prosodic information is necessary for listeners to be able to identify the affective tone of the utterance. At lengths below this, there was not enough information for listeners to process the emotional context of the utterance.[32]

Cognitive, Neural, Developmental, and Clinical Aspects

Child language

Unique prosodic features have been noted in infant-directed speech (IDS) - also known as baby talk, child-directed speech (CDS), or "motherese". Adults, especially caregivers, speaking to young children tend to imitate childlike speech by using higher and more variable pitch, as well as an exaggerated stress. These prosodic characteristics are thought to assist children in acquiring phonemes, segmenting words, and recognizing phrasal boundaries. And though there is no evidence to indicate that infant-directed speech is necessary for language acquisition, these specific prosodic features have been observed in many different languages.[33]

Aprosodia

An aprosodia is an acquired or developmental impairment in comprehending or generating the emotion conveyed in spoken language. Aprosody is often accompanied by the inability to properly utilize variations in speech, particularly with deficits in the ability to accurately modulate pitch, loudness, intonation, and rhythm of word formation.[34] This is seen sometimes in autistic individuals.[35]

The three main types of aprosodia are:

  • Lexical prosody: aprosodia affecting certain stressed syllables. Studies have shown that in the brain, lexical-tone contrast evokes a stronger pre-attentive response in the right hemisphere than in the left hemisphere. With consonant contrast, the brain produced an opposite pattern, with a stronger pre-attentive response in the left hemisphere than in the right hemisphere. An example of lexical prosody would be "CONvert" versus "conVERT".
  • Phrasal prosody: aprosodia affecting certain stressed words. Deficits in the left hemisphere affect this linguistic rule. An example of phrasal prosody would be a "Hot DOG", a dog that’s hot, versus a "HOT dog", a frankfurter.
  • Clausal prosody: aprosodia affecting contrastive, emphatic, and focal stress. Deficits in the left hemisphere affect this linguistic rule. An example of clausal prosody would be, "the horses were racing from the BARN" versus "the HORSES were racing from the barn."

Lexical prosody

Lexical prosody refers to the specific amplitudes, pitches, or lengths of vowels that are applied to specific syllables in words based on what the speaker wants to emphasize. The different stressors placed on individual syllables can change entire meanings of a word. Take one popular English word for example:

  • CONvert (noun: someone who has changed beliefs)
  • conVERT (verb: the act of changing)

In English, lexical prosody is used for a few different reasons. As we have seen above, lexical prosody was used to change the form of a word from a noun to a verb. Another function of lexical prosody has to do with the grammatical role that a word plays within a sentence. Adjectives and nouns of a sentence are often stressed on the first syllables while verbs are often stressed on the second syllable. For example:

  • "Elizabeth felt an increase in her happiness after meeting Tom"

Here, adults will emphasize the first syllable, "IN", as "increase" functions as an adjective.

  • "Tom will increase his workload"

Here, adults will emphasize the second syllable, "CREASE", as "increase" functions as a verb.

Another way that lexical prosody is used in the English language is in compound nouns such as "wishbone, mailbox, and blackbird" where the first compound is emphasized. Some suffixes can also affect the ways in which different words are stressed. Take "active" for example. Without the suffix, the lexical emphasis is on "AC". However, when we add the suffix -ity, the stress shifts to "TIV".[36]

Phrasal prosody

Phrasal prosody refers to the rhythm and tempo of phrases, often in an artistic setting such as music or poetry, but not always. The rhythm of the English language has four different elements: stress, time, pause, and pitch. Furthermore, "When stress is the basis of the metric pattern, we have poetry; when pitch is the pattern basis, we have rhythmic prose" (Weeks 11). Stress retraction is a popular example of phrasal prosody in everyday life. For example:

  • After eating sevenTEEN, PICKLES did not scare him
  • After eating SEVENteen PICKLES, he was afraid

Contrastive stress is another everyday English example of phrasal prosody that helps us determine what parts of the sentence are important. Take these sentences for example:

  • A man went up the STAIRS

Emphasizing that the STAIRS is how the man went up.

  • A MAN went up the stairs

Emphasizing that it was a MAN who went up the stairs.[37]

It's important to note that the right hemisphere of the brain dominates one's perception of prosody. In contrast to left hemisphere damage where patterns of aphasias are present, patterns of aprosodias are present with damage to the left hemisphere. In patients with right hemisphere lesions, they are characterized as monotonous and as lacking variety in their tone and expression. They're also seen to struggle with the identification and discrimination of semantically neutral sentences with varying tones of happiness, sadness, anger, and indifference, exemplifying the importance of prosody in language comprehension and production.

Brain regions involved

Producing these nonverbal elements requires intact motor areas of the face, mouth, tongue, and throat. This area is associated with Brodmann areas 44 and 45 (Broca's area) of the left frontal lobe. Damage to areas 44/45, specifically on the right hemisphere, produces motor aprosodia, with the nonverbal elements of speech being disturbed (facial expression, tone, rhythm of voice).

Understanding these nonverbal elements requires an intact and properly functioning right-hemisphere perisylvian area, particularly Brodmann area 22 (not to be confused with the corresponding area in the left hemisphere, which contains Wernicke's area).[38] Damage to the right inferior frontal gyrus causes a diminished ability to convey emotion or emphasis by voice or gesture, and damage to right superior temporal gyrus causes problems comprehending emotion or emphasis in the voice or gestures of others. The right Brodmann area 22 aids in the interpretation of prosody, and damage causes sensory aprosodia, with the patient unable to comprehend changes in voice and body language.

See also

References

  1. ^ Jones, Daniel (2011). Roach, Peter; Setter, Jane; Esling, John (eds.). Cambridge English Pronouncing Dictionary (18th ed.). Cambridge University Press. ISBN 978-0-521-15255-6.
  2. ^ "prosody". Merriam-Webster.com Dictionary. Merriam-Webster. Retrieved 2021-03-07.
  3. ^ "prosody | literature". Encyclopedia Britannica. Retrieved 2021-06-28.
  4. ^ a b Hirst, D.; Di Cristo, A. (1998). Intonation systems. Cambridge. pp. 4–7.
  5. ^ See for example Lai, Li-Fang; Gooden, Shelome (2016). "Acoustic cues to prosodic boundaries in Yami: A first look". Proceedings of Speech Prosody 8, May 31 – June 3, Boston, USA.
  6. ^ a b Ackerman, Lauren; Boone, Haley; Martinez, Michal Temkin (2016). "The sounds of language: An introduction to phonetics and phonology by Elizabeth C. Zsiga". Language. 92 (4): e292–e295. doi:10.1353/lan.2016.0082. ISSN 1535-0665. S2CID 152046615.
  7. ^ O'Connor, J.D and G.F.Arnold (1973). The Intonation of Colloquial English. Longman.
  8. ^ Crystal, David (1969). Prosodic Systems and Intonation in English. Cambridge University Press. ISBN 978-0-521-07387-5.
  9. ^ Halliday, M.A.K. (1967). Intonation and grammar in British English. The Hague: Mouton.
  10. ^ J.C. Wells (2007). English intonation. CUP.
  11. ^ Collins, B.; Mees, I. (2013) [First published 2003]. Practical Phonetics and Phonology: A Resource Book for Students (3rd ed.). Routledge. p. 129. ISBN 978-0-415-50650-2.
  12. ^ Cruttenden, Alan (1997). Intonation (2nd ed.). Cambridge. p. 13.
  13. ^ Ashby, M.; Maidment, J. (2005). Introducing Phonetic Science. Cambridge. pp. 167–8.
  14. ^ Hirst, D.; Di Cristo, A. (1998). Intonation systems. Cambridge. pp. 1–13.
  15. ^ "powtut1-1". Sfs.uni-tuebingen.de. Retrieved 2019-04-09.
  16. ^ Cruttenden, A. (1997). Intonation (2nd ed.). Cambridge. pp. 68–125. ISBN 0-521-59825-7.
  17. ^ Wells, J. (2006). English Intonation. Cambridge. pp. 187–194.
  18. ^ Stoyneshka, I.; Fodor, J.; Férnandez, E. M. (April 7, 2010). "Phoneme restoration methods for investigating prosodic influences on syntactic processing". Language and Cognitive Processes. 25 (7–9): 1265–1293. doi:10.1080/01690961003661192. S2CID 144478809.
  19. ^ Carroll, David W. (1994). Psychology of Language. Brooks/Cole. p. 87. ISBN 9780534213008.
  20. ^ Aitchison, Jean (1994). Words in the Mind. Blackwell. pp. 136–9. ISBN 9780631189213.
  21. ^ Wells, John (2006). English Intonation. Cambridge. pp. 116–124.
  22. ^ Frank Rudzicz (14 March 2016). Clear Speech: Technologies that Enable the Expression and Reception of Language. Morgan & Claypool Publishers. pp. 25–. ISBN 978-1-62705-827-8.
  23. ^ Pronina, Mariia; Hübscher, Iris; Vilà-Giménez, I.; Prieto, Pilar (2021). "Bridging the gap between prosody and pragmatics: The acquisition of pragmatic prosody in the preschool years and its relation with Theory of Mind". Frontiers in Psychology. 12: 662124. doi:10.3389/fpsyg.2021.662124. PMC 8322765. PMID 34335373.
  24. ^ Skantze, Gabriel (2021). "Turn-taking in conversational systems and human-robot interaction: a review". Computer Speech & Language. 67: 101178. doi:10.1016/j.csl.2020.101178.
  25. ^ Cole, Jennifer (2015). "Prosody in context: A review". Language, Cognition and Neuroscience. 30 (1–2): 1–31. doi:10.1080/23273798.2014.963130.
  26. ^ Brazil, David; Coulthard, Malcolm; Johns, Catherine (1980). Discourse Intonation and Language Teaching. Longman.
  27. ^ Cheang, H.S.; Pell, M.D. (May 2008). "The sound of sarcasm". Speech Communication. 50 (5): 366–81. doi:10.1016/j.specom.2007.11.003. S2CID 13310068.
  28. ^ Charles Darwin (1871). "The Descent of Man". Archived from the original on 2008-03-11. citing Johann Rudolph Rengger, Natural History of the Mammals of Paraguay, s. 49
  29. ^ R. Barra; J.M. Montero; J. Macías-Guarasa; L.F. D'Haro; R. San-Segundo; R. Córdoba. "Prosodic and segmental rubrics in emotion identification" (PDF). Archived (PDF) from the original on 2007-08-12.
  30. ^ H.-N. Teodorescu and Silvia Monica Feraru. (2007). "A Study on Speech with Manifest Emotions". Text, Speech and Dialogue. Lecture Notes in Computer Science. Vol. 4629. pp. 254–261. doi:10.1007/978-3-540-74628-7_34. ISBN 978-3-540-74627-0.
  31. ^ J.Pittam and K.R. Scherer (1993). "Vocal Expression and Communication of Emotion", Handbook of Emotions, New York, New York: Guilford Press.
  32. ^ Pell, M. D. (2005). "Prosody–face Interactions in Emotional Processing as Revealed by the Facial Affect Decision Task". Journal of Nonverbal Behavior. 29 (4): 193–215. doi:10.1007/s10919-005-7720-z. S2CID 13817074.
  33. ^ Gleason, Jean Berko., and Nan Bernstein Ratner. "The Development of Language", 8th ed. Pearson, 2013.
  34. ^ Elsevier. (2009). Mosby's Medical Dictionary, 8th edition.
  35. ^ McPartland J, Klin A (2006). "Asperger's syndrome". Adolesc Med Clin. 17 (3): 771–88. doi:10.1016/j.admecli.2006.06.010 (inactive 2024-09-12). PMID 17030291.{{cite journal}}: CS1 maint: DOI inactive as of September 2024 (link)
  36. ^ Schwanenflugel, Paula J.; Benjamin, Rebekah George (2017-01-01). "Lexical prosody as an aspect of oral reading fluency". Reading and Writing. 30 (1). Springer Link: 143–162. doi:10.1007/s11145-016-9667-3. ISSN 1573-0905.
  37. ^ Weeks, Ruth Mary. "Phrasal Prosody". The English Journal, vol. 10, no. 1, 1921, pp. 11–19. JSTOR, doi:10.2307/802950. Accessed 11 Dec 2023.
  38. ^ Miller, Lisa A; Collins, Robert L; Kent, Thomas A (2008). "Language and the modulation of impulsive aggression". The Journal of Neuropsychiatry and Clinical Neurosciences. 20 (3): 261–73. doi:10.1176/appi.neuropsych.20.3.261. PMID 18806230.

Further reading

  • Nespor, Marina. Prosody: an interview with Marina Nespor ReVEL, vol. 8, n. 15, 2010.
  • Nolte, John. The Human Brain 6th Edition
  • Mathews, Mark H. "Psychological trauma - The Trauma of Voices and Body Mentalists". traumaofvoices.com. Archived from the original on 18 March 2018. Retrieved 7 May 2018.
  • Shapiro, Karl., Beum, Robert. The Prosody Handbook: A Guide to Poetic Form. United States: Dover Publications, 2012. ISBN 9780486122670
  • Fox, Anthony. Prosodic Features and Prosodic Structure: The Phonology of 'Suprasegmentals'. United Kingdom: OUP Oxford, 2000. ISBN 9780191589768
  • Romero-Trillo, Jesús. Pragmatics and Prosody in English Language Teaching. Germany: Springer, 2012. ISBN 9789400738829

Read other articles:

Provincial park in Ontario, Canada La Cloche Provincial ParkIUCN category II (national park)Riviere La ClocheLocationUnorganized SE Algoma, Sables-Spanish RiversNearest townMassey, OntarioCoordinates46°06′18″N 82°01′30″W / 46.105°N 82.025°W / 46.105; -82.025Area7,448 ha (28.76 sq mi)[1]Established1985Governing bodyOntario ParksWebsitewww.ontarioparks.com/park/lacloche La Cloche Provincial Park is a provincial park at the bounda…

C-C motif chemokine ligand 3 المعرفات الأسماء المستعارة PAT 464.1, SIS-beta, C-C motif chemokine 3, MIP-1-alpha, CCL3, chemokine (C-C motif) ligand 3, G0/G1 switch regulatory protein 19-1, tonsillar lymphocyte LD78 alpha protein, small inducible cytokine A3 (homologous to mouse Mip-1a), macrophage inflammatory protein 1-alpha, Small-inducible cytokine A3 معرفات خارجية نمط التعبير عن الحمض النووي الريبوزي المزيد من بيا…

此條目可参照英語維基百科相應條目来扩充。 (2022年5月31日)若您熟悉来源语言和主题,请协助参考外语维基百科扩充条目。请勿直接提交机械翻译,也不要翻译不可靠、低品质内容。依版权协议,译文需在编辑摘要注明来源,或于讨论页顶部标记{{Translated page}}标签。 2022年印第安納州聯邦眾議員選舉 ← 2020 2022年11月8日 2024 → 印第安納州聯邦眾議員全部9個議席 &#…

Bradley Barron Renfro (Knoxville, Tennessee, 25 Juli 1982 - Los Angeles, California, 15 Januari 2008) merupakan seorang aktor berkebangsaan Amerika Serikat. Ia dikenal karena perannya dalam The Client (1994) sebagai lawan main Susan Sarandon) sekaligus sebagai debut karier aktingnya. Setelah namanya melambung, ia bermain dalam Apt Pupil (1998), sebuah adaptasi dari novel Stephen King. Ia juga bermain bersama Scarlett Johansson dalam Ghost World (2001). Brad dikenal sebagai pecandu heroin dan oba…

Questa voce o sezione sull'argomento gruppi musicali italiani non cita le fonti necessarie o quelle presenti sono insufficienti. Puoi migliorare questa voce aggiungendo citazioni da fonti attendibili secondo le linee guida sull'uso delle fonti. Segui i suggerimenti del progetto di riferimento. Banda musicale dell'Arma dei CarabinieriStato Italia CittàRoma DirettoreMassimo Martinelli Organico strumentale102 membri, organico Vesselliano Repertoriosinfonico, bandistico, leggero Periodo a…

Matteo Fedele Nazionalità  Svizzera Altezza 185 cm Peso 75 kg Calcio Ruolo Centrocampista Squadra  Chindia Târgoviște CarrieraGiovanili 2000-2004 Azzurri 902004-2008 Losanna2008-2009 Lilla2009-2013 SionSquadre di club1 2012-2015 Sion26 (1)2015→  Grasshoppers12 (0)2015-2016 Carpi9 (0)2016-2017→  Bari25 (4)2017-2018 Foggia9 (0)2018→  R.C. Cesena14 (1)2018-2019→  U Craiova20 (2)[1]2019-2021 Valenciennes12 (0)20…

California newspaper East Bay TimesFront page of the East Bay TimesTypeDaily newspaperFormatBroadsheetOwner(s)Bay Area News Group (Digital First Media)PublisherSharon RyanEditorFrank PineFounded1947; 77 years ago (1947)Headquarters2121 N. California Blvd, Suite 290, Walnut Creek, California 94596 U.S.Circulation55,787 Daily100,200 Sunday (as of 2022)[1]ISSN2473-0351Websiteeastbaytimes.com The East Bay Times is a daily broadsheet newspaper based in Walnut Creek, Cali…

American football player and coach (born 1939) For other people named Bill Snyder, see Bill Snyder (disambiguation). Bill SnyderSnyder in 2009Current positionTitleSpecial ambassadorTeamKansas StateConferenceBig 12Biographical detailsBorn (1939-10-07) October 7, 1939 (age 84)St. Joseph, Missouri, U.S.Playing career1958Missouri1959–1962William Jewell Position(s)Quarterback, defensive backCoaching career (HC unless noted)1962Gallatin HS (MO) (assistant)1964–1965Indio HS (CA) (assistant)196…

Bog in Mačva, Serbia. ZasavicaZasavicaLocationMačva region, Vojvodina province (partially)Coordinates44°57′27″N 19°31′35″E / 44.957624°N 19.526369°E / 44.957624; 19.526369TypemarshPrimary inflowsZasavica RiverPrimary outflowsZasavica RiverBasin countriesSerbiaMax. length33 kilometres (21 mi)Max. width300 metres (330 yd)Surface area11.5 square kilometres (4.4 sq mi)Average depth2.5 metres (8.2 ft)Max. depth10 metres (33 ft)S…

البطولات الفرنسية 1938 - فردي السيدات جزء من البطولات الفرنسية 1938  البلد فرنسا  التاريخ 1938  الرياضة كرة المضرب  البطل(ة) سيموني ماثيو الوصيف(ة) نيلي لاندري النتيجة 6–0، 6–3 البطولات الفرنسية 1937 - فردي السيدات  البطولات الفرنسية 1939 - فردي السيدات  تعديل مصدري - تعديل…

2007 Russian legislative election ← 2003 2 December 2007 2011 → All 450 seats in the State Duma226 seats needed for a majorityTurnout63.71% Party Leader % Seats +/– United Russia Vladimir Putin 65.01 315 +92 Communist Gennady Zyuganov 11.70 57 +5 LDPR Vladimir Zhirinovsky 8.23 40 +4 SR Sergey Mironov 7.83 38 New This lists parties that won seats. See the complete results below.Winning party by region Chairman of the State Duma before Chairman of the State Duma after Boris…

Emperor of Japan (869–949) Emperor Yōzei陽成天皇(from the Ogura Hyakunin Isshu)Emperor of JapanReignDecember 18, 876 – March 4, 884CoronationJanuary 20, 877PredecessorSeiwaSuccessorKōkōBornJanuary 2, 869Somedono In, Heian Kyō (Kyōto)DiedOctober 23, 949(949-10-23) (aged 80)Heian Kyō (Kyōto)BurialKaguragaoka no Higashi no misasagi (神楽岡東陵) (Kyōto)Issue Prince Motoyoshi Prince Motonaga Prince Mototoshi Princess Chōshi Princess Genshi Prince Motohira Minamoto no Kiyoka…

Country house near Towcester, Northamptonshire, England Easton Neston House in 1987 Easton Neston undergoing building work in 2007 Easton Neston is a large grade I listed[1] country house in the parish of Easton Neston near Towcester in Northamptonshire, England. It was built by William Fermor, 1st Baron Leominster (1648–1711), in the Baroque style to the design of the architect Nicholas Hawksmoor.[2] Easton Neston is thought to be the only mansion which was solely the work of …

Pour les articles homonymes, voir Venizélos. Cet article est une ébauche concernant un homme politique grec. Vous pouvez partager vos connaissances en l’améliorant (comment ?) selon les recommandations des projets correspondants. Benizélos RoúphosΜπενιζέλος Ρούφος Biographie Date de naissance 1795 Lieu de naissance Patras, Empire ottoman Date de décès 18 mars 1868 Lieu de décès Patras, Royaume de Grèce Nationalité grecque Parti politique Parti français Conjoint…

سفارة دولة فلسطين لدى لبنان فلسطين لبنان البلد لبنان  المكان بيروت العنوان الجناح، شارع عدنان الحكيم الاختصاص لبنان  السَفير أشرف دبور (منذُ 1 يناير 2012) الموقع الالكتروني www.palembassy-lb.net// تعديل مصدري - تعديل   سفارة دولة فلسطين لدى لبنان هي الممثلية الدبلوماسية العُليا ل…

此條目缺少有關频率 (统计学)的信息。 (2024年5月29日)請擴充此條目相關信息。討論頁可能有詳細細節。 三個閃動的光圓,從最低頻率(上端)至最高頻率(下端)。 频率(frequency)又稱週率,是物理学上描述某具规律週期性的现象或事件,在每单位时间内(即每秒)重复发生的次数(週期數,即循环次数);通常以符号 f {\displaystyle f} 或 ν {\displaystyle \nu } 表示。频率的…

Robert A. Heinlein Heinlein dédicaçant un de ses livres en 1976. Données clés Nom de naissance Robert Anson Heinlein Alias Anson McDonaldLyle MonroeJohn RiversideCaleb SaundersSimon York Naissance 7 juillet 1907 Butler, Missouri, États-Unis Décès 8 mai 1988 (à 80 ans) Carmel, Californie, États-Unis Activité principale Romancier, nouvelliste,essayiste, scénariste Distinctions Prix HugoPrix Damon-Knight Memorial Grand MasterPrix LocusPrix PrometheusMédaille de la NASA Auteur Langu…

This article relies excessively on references to primary sources. Please improve this article by adding secondary or tertiary sources. Find sources: Ottawa Bluesfest – news · newspapers · books · scholar · JSTOR (May 2015) (Learn how and when to remove this message) Ottawa BluesfestAerial view of the 2007 BluesfestGenreHip Hop, Reggae, Blues, pop, EDM and rockDatesJuly (6-16, 2023) (10 days)Location(s)Ottawa, Ontario, CanadaYears active1994-2019, 2022-pre…

Biografi ini tidak memiliki referensi atau sumber sehingga isinya tidak dapat dipastikan. Bantu memperbaiki artikel ini dengan menambahkan sumber tepercaya. Materi kontroversial atau trivial yang sumbernya tidak memadai atau tidak bisa dipercaya harus segera dihapus.Cari sumber: Ashley Boettcher – berita · surat kabar · buku · cendekiawan · JSTOR (Pelajari cara dan kapan saatnya untuk menghapus pesan templat ini) Ashley BoettcherPekerjaanAktrisTahun …

Late 16C portrait of Richard III (National Portrait Gallery, London), copied from an early 16C one in the Royal Collections. This version features in Josephine Tey's novel The Daughter of Time.Richard III of England has been depicted in literature and popular culture many times. In the Tudor period he was invariably portrayed as a villain, most famously in Shakespeare's play Richard III, but also in other literature of the period. Richard's life was not much depicted again until the 20th century…