List of the Cenozoic life of Utah

This list of the Cenozoic life of Utah contains the various prehistoric life-forms whose fossilized remains have been reported from within the US state of Utah and are between 66 million and 10,000 years of age.

A

• Abies
• †Achaenodon
  • †Achaenodon uintensis – type locality for species
• †Acmeodon
• †Acrocera
  • †Acrocera hirsuta – type locality for species
• †Aenocyon
  • †Aenocyon dirus – or unidentified comparable form
• †Aepinacodon
• †Aethomylos
  • †Aethomylos simplicidens
• †Ageina – or unidentified comparable form

• 

  Life restoration of the Eocene-Oligocene even-toed ungulate Agriochoerus

  †Agriochoerus
  • †Agriochoerus maximus
• Alligator
• †Allognathosuchus – or unidentified comparable form
• †Allophaiomys
  • †Allophaiomys pliocaenicus
• Amia
  • †Amia uintensis
• Amyda
  • †Amyda crassa – type locality for species
  • †Amyda gregaria
• †Amynodon
  • †Amynodon advenus
  • †Amynodon reedi
• †Anaptomorphus
  • †Anaptomorphus aemulus
• †Anemorhysis
• †Anisonchus
  • †Anisonchus athelas
  • †Anisonchus oligistus
  • †Anisonchus onostus
  • †Anisonchus sectorius

• 

  Fossilized shell of the Eocene-Oligocene turtle Anosteira

  †Anosteira
  • †Anosteira ornata
• Anthonomus
  • †Anthonomus soporus
• †Antiacodon
  • †Antiacodon pygmaeus
• Antilocapra
  • †Antilocapra americana – or unidentified comparable form
• Apalone
• †Apataelurus
  • †Apataelurus kayi – type locality for species
• †Apatemys
  • †Apatemys bellulus – or unidentified comparable form
  • †Apatemys bellus – or unidentified comparable form
  • †Apatemys rodens – or unidentified comparable form
  • †Apatemys uintensis – type locality for species
• †Aphronorus
  • †Aphronorus simpsoni

• 

  Restoration of an Arctodus, or short-faced bear, with a human to scale

  †Arctodus
  • †Arctodus simus
• †Armintodelphys
  • †Armintodelphys dawsoni
• †Artemisia
• †Asilopsis – type locality for genus
  • †Asilopsis fusculus – type locality for species
• †Asilus
  • †Asilus palaeolestes – type locality for species
• †Aulobaris
  • †Aulobaris circumscripta
  • †Aulobaris comminuta – type locality for species
• †Auxontodon
  • †Auxontodon pattersoni

B

• †Baena
  • †Baena arenosa – type locality for species
  • †Baena emiliae
• †Baptemys
• †Basirepomys
  • †Basirepomys robertsi – type locality for species
• †Bembidium
  • †Bembidium exoletum – type locality for species
• Bison
  • †Bison latifrons – tentative report
• †Bittacus
  • †Bittacus egestionis – type locality for species
• †Blickomylus
  • †Blickomylus moroni – type locality for species
• †Bootherium
  • †Bootherium bombifrons

• 

  Fossilized skeleton of the Eocene crocodilian Boverisuchus

  †Boverisuchus
  • †Boverisuchus vorax – type locality for species
• †Brachyhyops
  • †Brachyhyops wyomingensis
• Brachylagus
  • †Brachylagus idahoensis – or unidentified comparable form
• †Bridgeremys
  • †Bridgeremys pusilla – tentative report
• †Bunomeryx
  • †Bunomeryx montanus – type locality for species

C

• 

  Life restoration of the Pliocene-Holocene camel Camelops

  †Camelops
  • †Camelops hesternus – or unidentified comparable form
• Camponotus
  • †Camponotus vetus – type locality for species
• Canis
  • †Canis latrans – tentative report
  • †Canis lupus
• †Capricamelus
• †Carabites
  • †Carabites eocenicus – type locality for species
  • †Carabites exanimus – type locality for species
• †Catopsalis
  • †Catopsalis fissidens
• †Cedrelospermum
  • †Cedrelospermum nervosum
• †Centetodon
  • †Centetodon bembicophagus
  • †Centetodon pulcher

• 

  Life restoration of the Late Cretaceous-Eocene choristoderan reptile Champsosaurus

  †Champsosaurus
• Chara
• Cheilosia
  • †Cheilosia scudderi
• †Chipetaia
  • †Chipetaia lamporea – type locality for species
• †Chisternon
  • †Chisternon undatum – type locality for species
• †Chriacus
  • †Chriacus baldwini
• †Cicindelopsis – type locality for genus
  • †Cicindelopsis eophilus – type locality for species
• †Colodon
• †Compsemys
  • †Compsemys victa
• †Conacodon
  • †Conacodon kohlbergeri
• †Coniatus
  • †Coniatus refractus
• †Conoryctella – type locality for genus
  • †Conoryctella dragonensis – type locality for species
  • †Conoryctella pattersoni
• †Copemys
  • †Copemys mariae – or unidentified related form
• †Coryphodon
  • †Coryphodon lobatus
• †Cristadjidaumo – type locality for genus
  • †Cristadjidaumo mckennai – type locality for species
• Crocodylus

  • 

    Fossilized skull of the Eocene crocodilian Crocodylus acer

    †Crocodylus acer – type locality for species
  • †Crocodylus affinis
• †Crypholestes
• Culex
  • †Culex proavitus – type locality for species
• †Cupidinimus
• †Cuterebra
  • †Cuterebra ascarides
• †Cyttaromyia – type locality for genus
  • †Cyttaromyia fenestrata – type locality for species

D

• †Desmatoclaenus
  • †Desmatoclaenus hermaeus
• Dicranomyia
  • †Dicranomyia primitiva – type locality for species
  • †Dicranomyia rostrata – type locality for species
  • †Dicranomyia stigmosa – type locality for species

• 

  Mounted fossilized skeleton of the Miocene-Pliocene horse Dinohippus

  †Dinohippus
• †Diplacodon
  • †Diplacodon elatus
  • †Diplacodon emarginatus – type locality for species
• †Diplobunops
  • †Diplobunops crassus
  • †Diplobunops matthewi
• †Diprionomys
  • †Diprionomys minimus – or unidentified comparable form
• †Dipsalidictis
  • †Dipsalidictis transiens
• †Draconodus
  • †Draconodus apertus
• †Dracontolestes
  • †Dracontolestes aphantus – type locality for species
• †Duchesnehippus
  • †Duchesnehippus intermedius
• †Duchesneodus
  • †Duchesneodus uintensis

E

• 

  Fossilized shell of the Eocene turtle Echmatemys

  †Echmatemys
  • †Echmatemys depressa – type locality for species
  • †Echmatemys douglassi – type locality for species
  • †Echmatemys septaria – type locality for species
  • †Echmatemys uintensis
• †Ectoconus
  • †Ectoconus ditrigonus
  • †Ectoconus symbolus
• †Ellipsodon
  • †Ellipsodon grangeri
  • †Ellipsodon lemuroides
  • †Ellipsodon sternbergi – type locality for species
• †Elliptio

• 

  Life restoration of a pair of the Eocene uintathere mammal Eobasileus. Charles R. Knight (1890s).

  †Eobasileus
  • †Eobasileus cornutus
• †Eobracon – type locality for genus
  • †Eobracon cladurus – type locality for species
• †Eocaenonemonyx
  • †Eocaenonemonyx kuscheli – type locality for species
• †Eoconodon
• †Eodiploglossus
  • †Eodiploglossus borealis
• †Eoformica
  • †Eoformica pinguis – type locality for species
• †Eomoropus
  • †Eomoropus amarorum – type locality for species
  • †Eomoropus anarsius
• †Eonessa – type locality for genus
  • †Eonessa anaticula – type locality for species
• †Epicaerus
  • †Epicaerus effossus
  • †Epicaerus eradicatus
  • †Epicaerus evigoratus
  • †Epicaerus exanimis
  • †Epicaerus subterraneus
  • †Epicaerus terrosus
• †Epihippus
  • †Epihippus gracilis
• †Epiphanis
  • †Epiphanis deletus – type locality for species
• †Epitriplopus
  • †Epitriplopus uintensis
• Equus
  • †Equus conversidens – tentative report
• Erethizon
  • †Erethizon dorsatum – tentative report
• †Eucastor
  • †Eucastor tortus – or unidentified comparable form

• 

  Fossil of the Eocene bee Euglossopteryx

  †Euglossopteryx – type locality for genus
  • †Euglossopteryx biesmeijeri – type locality for species
• Eutamias
  • †Eutamias minimus

G

• †Glyptosaurus
  • †Glyptosaurus sylvestris
• †Gnoriste
  • †Gnoriste dentoni – type locality for species
• †Goniacodon
  • †Goniacodon hiawathae
• †Griphomys

H

Life restoration of the Eocene horse Eohippus. Heinrich Harder (1920).

• †Hadrianus
  • †Hadrianus corsoni
  • †Hadrianus utahensis – type locality for species
• †Haplaletes
  • †Haplaletes andakupensis
• †Haploconus
  • †Haploconus angustus
  • †Haploconus elachistus
• †Haplolambda – or unidentified comparable form
  • †Haplolambda simpsoni – type locality for species

• 

  Fossilized skull of the mesonychian mammal Harpagolestes

  †Harpagolestes
  • †Harpagolestes brevipes – type locality for species
  • †Harpagolestes leotensis – type locality for species
  • †Harpagolestes uintensis
• †Helaletes
  • †Helaletes nanus
• †Helodermoides – tentative report
• †Heptacodon
• †Herpetotherium
  • †Herpetotherium innominatum
• †Hesperolagomys
  • †Hesperolagomys galbreathi
• †Hessolestes
  • †Hessolestes ultimus – type locality for species
• †Heteraletes
  • †Heteraletes leotanus
• †Hoplochelys

• 

  Life restoration of the Eocene-Miocene creodont mammal Hyaenodon

  †Hyaenodon
  • †Hyaenodon vetus – or unidentified comparable form
• Hydrobia – or unidentified comparable form
• †Hylobius
  • †Hylobius deleticius – type locality for species
• †Hylomeryx
  • †Hylomeryx annectens
  • †Hylomeryx quadricuspis – type locality for species
• †Hyopsodus
  • †Hyopsodus minusculus
  • †Hyopsodus paulus
  • †Hyopsodus uintensis – type locality for species
• †Hypohippus
• †Hypolagus
  • †Hypolagus vetus
• †Hyrachyus
  • †Hyrachyus eximius
  • †Hyrachyus modestus
• †Hyracodon
  • †Hyracodon medius
• †Hyracotherium
  • †Hyracotherium vasacciense

I

• †Isectolophus – type locality for genus
  • †Isectolophus annectens – type locality for species

J

• †Janimus
  • †Janimus rhinophilus – type locality for species

K

• †Kimbetohia
  • †Kimbetohia campi

L

• Laccophilus
• †Lambdotherium
• †Lasioptera
  • †Lasioptera recessa – type locality for species
• †Leistotrophus
  • †Leistotrophus patriarchicus – type locality for species
• Lemmiscus
  • †Lemmiscus curtatus
• Lepisosteus
  • †Lepisosteus simplex

• 

  Life restoration of the Eocene-Oligocene even-toed ungulate Leptomeryx

  †Leptomeryx
• †Leptoreodon
  • †Leptoreodon marshi
• †Leptotragulus
  • †Leptotragulus clarki
  • †Leptotragulus medius – type locality for species
  • †Leptotragulus proavus
• Lepus
  • †Lepus townsendii – or unidentified comparable form
• †Limalophus
  • †Limalophus compositus
  • †Limalophus receptus – type locality for species

• 

  Top view of a fossilized skull of the Eocene creodont mammal Limnocyon

  †Limnocyon
  • †Limnocyon potens
  • †Limnocyon verus – or unidentified comparable form
• †Liodontia
• †Lioplacoides – or unidentified comparable form
• †Litaletes
  • †Litaletes gazini
• †Lithadothrips – type locality for genus
  • †Lithadothrips vetusta – type locality for species
• †Loxolophus
  • †Loxolophus pentacus
  • †Loxolophus spiekeri

• 

  A living Lynx

  Lynx
  • †Lynx canadensis – or unidentified comparable form

M

• †Macrotarsius
  • †Macrotarsius jepseni – type locality for species
• †Mammut
  • †Mammut americanum
• †Mammuthus
  • †Mammuthus columbi

  • 

    Life restorations of a Mammut americanum, or American mastodon (right), and a Mammuthus primigenius, or wooly mammoth (left)

    †Mammuthus primigenius
• Marsilea[1]
  • †Marsilea sprungerorum[1]
• †Megalamynodon
  • †Megalamynodon regalis – type locality for species
• †Megalonyx
  • †Megalonyx jeffersonii
• †Menops
  • †Menops marshi
• †Merriamoceros
• †Mesomeryx – type locality for genus
  • †Mesomeryx grangeri – type locality for species

• 

  Life restoration of the Eocene mammal Mesonyx

  †Mesonyx
  • †Mesonyx obtusidens
• †Metaliomys – type locality for genus
  • †Metaliomys sevierensis – type locality for species
• †Metanoiamys
  • †Metanoiamys lacus
• †Metarhinus
  • †Metarhinus diploconus
  • †Metarhinus fluviatilis – type locality for species
  • †Metarhinus parvus
• †Metatelmatherium
  • †Metatelmatherium ultimum – type locality for species

• 

  Life restoration of the Paleocene-Eocene mammal Miacis

  †Miacis
  • †Miacis parvivorus – or unidentified comparable form
• †Microparamys
  • †Microparamys dubius
  • †Microparamys minutus
• †Micropternodus
• †Microsyops
• Microtus
  • †Microtus montanus
  • †Microtus paroperarius
• Mictomys
  • †Mictomys borealis
• †Mimomys
  • †Mimomys dakotaensis – or unidentified comparable form
• †Mimotricentes
• †Miocyon
  • †Miocyon major – or unidentified comparable form
  • †Miocyon scotti – type locality for species
  • †Miocyon vallisrubrae – type locality for species
• Mustela
  • †Mustela richardsonii – tentative report
• †Myrmecoboides
• †Mytonolagus
  • †Mytonolagus petersoni – type locality for species
• †Mytonomeryx
  • †Mytonomeryx scotti – type locality for species
• †Mytonomys
  • †Mytonomys mytonensis
  • †Mytonomys robustus – type locality for species

N

A living Neogale vison, or American mink

• †Nemotelus
  • †Nemotelus eocenicus – type locality for species
• Neogale
  • †Neogale vison
• †Neoplagiaulax
  • †Neoplagiaulax macintyrei
• Neoptochus – tentative report
• Neotamias
• Neotoma
• †Notharctus

  • 

    Mounted fossilized skeleton of the Eocene primate Notharctus tenebrosus

    †Notharctus tenebrosus – or unidentified comparable form
• †Nyctitherium
  • †Nyctitherium serotinum

O

• 

  A living Odocoileus deer

  Odocoileus
• †Oliveremys
  • †Oliveremys uintaensis
• †Omomys
  • †Omomys carteri
  • †Omomys lloydi – type locality for species
• Ondatra
• †Onychodectes
  • †Onychodectes tisonensis – or unidentified comparable form
• †Oodectes – tentative report
• Ophryastes
  • †Ophryastes petrarum
• †Ophryastites
  • †Ophryastites digressus – type locality for species
• †Orelladjidaumo – or unidentified comparable form
• †Oromeryx
  • †Oromeryx plicatus – type locality for species
• †Ourayia
  • †Ourayia hopsoni
  • †Ourayia uintensis – type locality for species
• †Oxyacodon
  • †Oxyacodon apiculatus
  • †Oxyacodon ferronensis – type locality for species
  • †Oxyacodon marshater – type locality for species
• †Oxyaenodon
  • †Oxyaenodon dysodus
• †Oxyclaenus
  • †Oxyclaenus pugnax
• †Oxygonus
  • †Oxygonus mortuus – type locality for species
• †Oxytomodon
  • †Oxytomodon perissum

P

• †Pachymerus
  • †Pachymerus petrensis – type locality for species
• †Paenemarmota
  • †Paenemarmota sawrockensis
• †Palaeictops
  • †Palaeictops bridgeri
• †Palaeoeristalis – type locality for genus
  • †Palaeoeristalis tesselatus – type locality for species
• †Palaeothrips – type locality for genus
  • †Palaeothrips fossilis – type locality for species
• †Palaeoxantusia
• †Pantolestes
  • †Pantolestes longieundus – or unidentified comparable form
• †Paradjidaumo
  • †Paradjidaumo alberti – or unidentified comparable form
  • †Paradjidaumo trilophus

• 

  Fossilized skeleton of the Pliocene-Pleistocene ground sloth Paramylodon

  †Paramylodon
  • †Paramylodon harlani – or unidentified comparable form
• †Paramys
  • †Paramys compressidens – type locality for species
  • †Paramys delicatus
  • †Paramys leptodus
• †Parasauromalus
  • †Parasauromalus olseni
• †Parectypodus
• †Pareumys
  • †Pareumys grangeri
  • †Pareumys guensburgi – type locality for species
  • †Pareumys milleri – type locality for species
  • †Pareumys troxelli
• †Paromomys
  • †Paromomys depressidens
• †Paronychomys
  • †Paronychomys lemredfieldi
• †Paropsocus – type locality for genus
  • †Paropsocus disjunctus – type locality for species
• †Passaliscomys – type locality for genus
  • †Passaliscomys priscus – type locality for species
• †Pauromys
• †Penetrigonias
  • †Penetrigonias dakotensis
• †Pentacemylus
  • †Pentacemylus leotensis
  • †Pentacemylus progressus – type locality for species
• †Peradectes
  • †Peradectes chesteri

• 

  Fossilized mandible of the Eocene-Miocene mammal Peratherium

  †Peratherium
  • †Peratherium marsupium
• †Periptychus
  • †Periptychus carinidens
  • †Periptychus coarctatus
• Peromyscus
  • †Peromyscus maniculatus
• Phenacomys
  • †Phenacomys gryci – or unidentified comparable form
  • †Phenacomys intermedius
• †Phygadeuon
  • †Phygadeuon petrifactellus – type locality for species
• Phyllobius
  • †Phyllobius avus
  • †Phyllobius carcerarius
• †Phyllophaga
  • †Phyllophaga avus – type locality for species
• Physa
• Picea

• 

  A living Pinus, or pine tree

  Pinus
• †Plastomenoides
  • †Plastomenoides lamberti
• †Plastomenus
• Plecia
  • †Plecia woodruffi – type locality for species
• †Plesiolestes
  • †Plesiolestes nacimienti
• †Poabromylus
  • †Poabromylus kayi – type locality for species
• †Poebrodon
  • †Poebrodon kayi – type locality for species
• †Polemonium – or unidentified comparable form
• †Presbyorniformipes
  • †Presbyorniformipes feduccii – type locality for species

• 

  Life restoration of the Paleocene-Eocene waterfowl Presbyornis

  †Presbyornis – type locality for genus
  • †Presbyornis pervetus – type locality for species
  • †Presbyornis recurvirostrus – type locality for species
• †Procaimanoidea – type locality for genus
  • †Procaimanoidea kayi
  • †Procaimanoidea utahensis – type locality for species
• †Procas
  • †Procas vinculatus
• †Procynodictis
  • †Procynodictis vulpiceps
• †Prodaphaenus
  • †Prodaphaenus uintensis – type locality for species
• †Promioclaenus
• †Pronophlebia – type locality for genus
  • †Pronophlebia rediviva – type locality for species
• †Propalaeosinopa – tentative report
• †Protadjidaumo
  • †Protadjidaumo typus – type locality for species
• †Protictis
  • †Protictis haydenianus

• 

  Restorative models in multiple views of the Eocene brontothere Protitanotherium (figure 3)

  †Protitanotherium
• †Protochriacus
  • †Protochriacus simplex
• †Protoptychus
  • †Protoptychus hatcheri
• †Protoreodon
  • †Protoreodon minor – type locality for species
  • †Protoreodon parvus
  • †Protoreodon petersoni – type locality for species
  • †Protoreodon pumilus – type locality for species
• †Protoselene
  • †Protoselene griphus

• 

  Illustration of a fossilized skull of the Eocene camel Protylopus

  †Protylopus
  • †Protylopus annectens – type locality for species
  • †Protylopus petersoni

• 

  Fossilized skeleton of the Eocene creodont mammal Proviverra

  †Proviverra
  • †Proviverra longipes
• †Proviverroides
  • †Proviverroides piercei – or unidentified comparable form
• †Proxestops – or unidentified comparable form
  • †Proxestops silberlingii
• †Pseudodiplacodon
• †Pseudosalix – type locality for genus
  • †Pseudosalix handleyi – type locality for species
• †Pseudotomus
  • †Pseudotomus eugenei – type locality for species
  • †Pseudotomus petersoni – type locality for species

• 

  Illustration of a fossilized skull of the Paleocene multituberculate mammal Ptilodus

  †Ptilodus
  • †Ptilodus mediaevus
  • †Ptilodus tsosiensis – or unidentified comparable form

R

• †Reithroparamys
  • †Reithroparamys sciuroides – type locality for species

• 

  A living Rhinoclemmys, or Neotropical wood turtle

  Rhinoclemmys – report made of unidentified related form or using admittedly obsolete nomenclature

S

• 

  Fossilized skeleton of the Eocene monitor lizard Saniwa

  †Saniwa
  • †Saniwa ensidens
• †Sargus
  • †Sargus vetus – type locality for species
• †Scenopagus
  • †Scenopagus edenensis
  • †Scenopagus priscus
• †Sciuravus
  • †Sciuravus altidens – type locality for species
  • †Sciuravus eucristadens
  • †Sciuravus popi – type locality for species
• †Simidectes – type locality for genus
  • †Simidectes magnus – type locality for species
  • †Simidectes medius – type locality for species
• †Simimeryx
  • †Simimeryx minutus
• †Simimys – or unidentified comparable form
• †Sinopa
  • †Sinopa agilis
  • †Sinopa major
  • †Sinopa rapax
• †Smilodectes
  • †Smilodectes gracilis

• 

  Life restoration of the Pleistocene-Holocene saber-tooth cat Smilodon

  †Smilodon
  • †Smilodon fatalis – or unidentified comparable form
• Sorex
  • †Sorex palustris
• Spermophilus
  • †Spermophilus armatus – or unidentified comparable form
• †Sphenocoelus
  • †Sphenocoelus hyognathus
• †Sthenodectes
  • †Sthenodectes incisivum – type locality for species
• †Stygimys
  • †Stygimys kuszmauli
• †Stylemys
  • †Stylemys uintensis – type locality for species

• 

  Life restoration of the Eocene taeniodont mammal Stylinodon mirus

  †Stylinodon
  • †Stylinodon mirus
• †Syntomostylus
  • †Syntomostylus rudis

T

• 

  Restoration of the Paleocene multituberculate mammal Taeniolabis

  †Taeniolabis
  • †Taeniolabis taoensis
• †Talpavus
  • †Talpavus duplus – type locality for species
  • †Talpavus nitidus
• †Tapocyon
  • †Tapocyon robustus
• Taxidea
  • †Taxidea taxus
• †Thisbemys
  • †Thisbemys medius
  • †Thisbemys uintensis – type locality for species
• Thomomys
  • †Thomomys talpoides
• †Tilgidopsis – type locality for genus
  • †Tilgidopsis haesitans – type locality for species
• Tinosaurus
  • †Tinosaurus stenodon
• †Tricentes
  • †Tricentes subtrigonus – type locality for species
• †Triplopus
  • †Triplopus implicatus
  • †Triplopus obliquidens
  • †Triplopus rhinocerinus – type locality for species
• †Trogosus
• †Tylocephalonyx

U

• †Uintaceras
  • †Uintaceras radinskyi – type locality for species
• †Uintacyon
  • †Uintacyon acutus – type locality for species
• †Uintanius
  • †Uintanius ameghini
• †Uintasorex
  • †Uintasorex parvulus – or unidentified comparable form

• 

  Life restoration of the Eocene mammal Uintatherium

  †Uintatherium
• †Utahia
  • †Utahia kayi – type locality for species

V

• †Valenia
  • †Valenia wilsoni
• †Viverravus
  • †Viverravus gracilis
  • †Viverravus minutus
  • †Viverravus sicarius
• Viviparus
• Vulpes

  • 

    A living Vulpes vulpes, or red fox

    †Vulpes vulpes

X

• †Xestops
  • †Xestops vagans
• †Xylotosyrphus – type locality for genus
  • †Xylotosyrphus pulchrafenestra – type locality for species

Z

• 

  A living Zapus jumping mouse

  Zapus
• †Zionodon – type locality for genus
  • †Zionodon satanus – type locality for species
  • †Zionodon walshi – type locality for species

References

1. Elizabeth J. Hermsen (2019). "Revisions to the fossil sporophyte record of Marsilea". Acta Palaeobotanica. 59 (1): 27–50. doi:10.2478/acpa-2019-0005.

• Various Contributors to the Paleobiology Database. "Fossilworks: Gateway to the Paleobiology Database". Retrieved 7 August 2017.