List of oldest trees

Pando, a colony of quaking aspen, is one of the oldest-known clonal trees. Recent estimates of its age range up to 14,000 years old, and 18,000 years by the latest (2024) estimate.[1] It is located in Utah, United States.

This is a list of the oldest-known trees, as reported in reliable sources. Definitions of what constitutes an individual tree vary. In addition, tree ages are derived from a variety of sources, including documented "tree-ring" (dendrochronological) count core samples, and from estimates. For these reasons, this article presents three lists of "oldest trees," each using varying criteria.

There are three tables of trees, which are listed by age and species. The first table includes trees for which a minimum age has been directly determined, either through counting or cross-referencing tree rings or through radiocarbon dating. Many of these trees may be even older than their listed ages, but the oldest wood in the tree has rotted away. For some old trees, so much of the center is missing that their age cannot be directly determined. Instead, estimates are made based on the tree's size and presumed growth rate. The second table includes trees with these estimated ages. The last table lists clonal colonies in which no individual tree trunks may be remarkably old but in which the organism as a whole is thought to be very old.

The current record-holders for individual, non-clonal trees are the Great Basin bristlecone pine trees from California and Nevada, in the United States. Through tree-ring cross-referencing, they have been shown to be almost five millennia old.[2]

A clonal colony can survive for much longer than an individual tree. A colony of 48,000 quaking aspen trees (nicknamed Pando), covering 106 acres (43 ha) in the Fishlake National Forest of Utah, is considered one of the oldest and largest organisms in the world. Recent estimates set the colony's age at several thousand (up to 14,000) years, although tree ring samples date individual stems at rarely more than 130 years.[3][4][5][6][7] A colony of Huon pine trees covering 2.5 acres (1.0 ha) on Mount Read (Tasmania) is estimated to be around 10,000 years old, as determined by DNA samples taken from pollen collected from the sediment of a nearby lake. Individual trees in this group date to no more than 4,000 years old, as determined by tree ring samples.[8]

Trees with verified ages

Name	Age (years)	Year Germinated	Species	Location	Country	Notes
Prometheus (WPN-114)	4,900+	2,900+ BCE	Great Basin bristlecone pine Pinus longaeva	Wheeler Peak, Nevada	United States	Cut down by Donald Rusk Currey in 1964.[9]
Methuselah	4,856[nb 1]	2,832 BCE	Great Basin bristlecone pine Pinus longaeva	White Mountains (California)	United States	It is the oldest known living (non-clonal) tree in the world.[11]
Alerce Milenario or Gran Abuelo	3,653–5,484	1,630+ BCE	Patagonian cypress Fitzroya cupressoides	Cordillera Pelada, Los Ríos	Chile	Alive.[11][12] New unconfirmed estimation of 5,484 years would make it the oldest (non-clonal) tree in the world.[13] Located within Alerce Costero National Park.[14]
CBR26	3,266	1,245 BCE	Giant sequoia Sequoiadendron giganteum	Sierra Nevada, California	United States	Dead.[11]
D-21	3,220	1,199 BCE	Giant sequoia Sequoiadendron giganteum	Sierra Nevada, California	United States	Dead.[11]
D-23	3,075	1,054 BCE	Giant sequoia Sequoiadendron giganteum	Sierra Nevada, California	United States	Dead.[11]
CMC 3	3,033	1,012 BCE	Giant sequoia Sequoiadendron giganteum	Sierra Nevada, California	United States	Dead.[11]
Scofield Juniper	2,675	654 BCE	Sierra juniper Juniperus grandis[15]	Sierra Nevada, California	United States	Dead.[11]
BLK227	2,649	626 BCE	Bald cypress Taxodium distichum	Three Sisters Cove, Black River (North Carolina)	United States	[16]
CB-90-11	2,467	443 BCE	Rocky Mountain bristlecone pine Pinus aristata	Central Colorado	United States	[11][17]
Panke Baobab	2,419	398 BCE	African baobab Adansonia digitata	Matabeleland North	Zimbabwe	Died in 2011; the oldest non-clonal angiosperm ever documented.[18]
Jaya Sri Maha Bodhi	2,311	287 BCE	Sacred fig Ficus religiosa	Anuradhapura, North Central Province	Sri Lanka	A sapling claimed to be from the Bodhi tree, the legendary tree under which the Buddha is said to have become enlightened. It was brought to its current location and planted at around 288 BCE by Sanghamitra, daughter of Emperor Ashoka. It is the oldest living human-planted tree in the world with a known planting date.[19]
?	2,230	209 BCE	Qilian juniper Juniperus przewalskii	Delingha, Qinghai Province	China	[11]
?	2,200	179 BCE	Coast redwood Sequoia sempervirens	Northern California	United States	Dead.[11]
Bennett Juniper	2,200	179 BCE	Sierra juniper Juniperus grandis[15]	Sierra Nevada, California	United States	[11]
SHP 7	2,110	89 BCE	Foxtail pine Pinus balfouriana	Sierra Nevada California	United States	[11]
BLK232	2,089	68 BCE	Bald cypress Taxodium distichum	Three Sisters Cove, Black River (North Carolina)	United States	[16]
?	1,951	74	Subalpine larch Larix lyallii	Kananaskis, Alberta	Canada	[11][20]
CRE 175	1,889	133	Rocky Mountain juniper Juniperus scopulorum	Northern New Mexico	United States	[11]
Miles Juniper	1,836	189	Sierra juniper Juniperus grandis^[citation needed]	Sierra Nevada, California	United States	[11][21]
Jōmon Sugi	1,813	212	Sugi Cryptomeria japonica	Yakushima Island	Japan	[22]
KET 3996	1,727	298	Limber pine Pinus flexilis	Ketchum, Idaho	United States	[11][23]
BFR-46	1,697	325	Limber pine Pinus flexilis	Wasatch Mountains, Utah	United States	[11]
FL117	1,686	339	Northern whitecedar Thuja occidentalis	Ontario	Canada	[11][24]
ERE	1,670	352	Limber pine Pinus flexilis	Northern New Mexico	United States	[11]
RCR 1	1,666	356	Foxtail pine Pinus balfouriana	Sierra Nevada, California	United States	[11]
?	1,661	361	Limber pine Pinus flexilis	South Park (Park County, Colorado)	United States	[11]
BBL 2	1,649	373	Foxtail pine Pinus balfouriana	Sierra Nevada, California	United States	[11]
BCK 69	1,658	376	Bald cypress Taxodium distichum	Bladen County, North Carolina	United States	[11][25]
?	1,636	386	Nootka cypress Callitropsis nootkatensis	Vancouver Island	Canada	[11]
FL101	1,567	455	Northern whitecedar Thuja occidentalis	Ontario	Canada	[11]
?	1,542	480	Limber pine Pinus flexilis	Central Colorado	United States	[11]
Italus	1,237	788	Heldreich's pine Pinus heldreichii	Pollino National Park	Italy	Alive. Oldest tree in Europe with verified age (radiocarbon calibrated tree-ring procedure).[26]

Trees with estimated ages

Note: The ages of the trees in this list are speculative and only serve as estimates.

Name	Age (years)	Species	Location	Country	Notes
Old Tjikko	9,565 (years)	Norway spruce Picea abies	Fulufjället, Dalarna	Sweden
Alerce Milenario or Gran Abuelo	5,484	Patagonian cypress Fitzroya cupressoides }	Cordillera Pelada, Los Ríos	Chile	A new 2022 estimation of 5,484 years expands on a previous minimum age based on incomplete tree rings of 3,653.[13][14]
?[nb 2]	5,076	Great Basin bristlecone pine Pinus longaeva	White Mountains (California)	United States	Tree cored by Edmund Schulman, age determined by Tom Harlan. However, core is missing and date is unconfirmed.[11][27]
Llangernyw Yew	4,000–5,000	Common yew Taxus baccata	Llangernyw, Conwy	United Kingdom	Girth of 10.75 m. Situated in the churchyard of St Dygain's Church in Llangernyw village, Wales. One of the 50 Great British Trees.
Sarv-e Abarkuh	4,500	Mediterranean cypress Cupressus sempervirens	Abarkuh, Yazd	Iran	Also called "Zoroastrian Sarv".[28][29]
Gümeli Porsuğu	4,115	Yew Taxus baccata	Zonguldak	Turkey	Alive, found in 2016.[30][31]
The Ancient Yew	4,000	Yew Taxus baccata	Tisbury, Wiltshire	United Kingdom	37 feet in circumference. Situated in the churchyard of St John's, Tisbury. Carbon dated by David Bellamy.[32]
The Senator	3,500	Pond cypress Taxodium ascendens	Longwood, Florida	United States	Died in January 2012 from human-caused fire.[33][34]
Oliveira do Mouchão (KNJ1/601)	3,350	European olive tree Olea europea L. var. europaea	Mouriscas, Abrantes	Portugal	Alive.[35][36]
S'Ozzastru	3,000–4,000	Olive Olea europaea	Luras, Sardinia	Italy	[37]
The President	3,200	Giant sequoia Sequoiadendron giganteum	Sierra Nevada, California	United States	Alive.
Fortingall Yew	3,000	Common yew Taxus baccata	Fortingall, Perthshire	United Kingdom	Alive. Possibly the oldest tree in Britain.[38][39]^[clarification needed]
Alishan Sacred Tree	3,000	Formosan cypress Chamaecyparis formosensis	Alishan National Scenic Area, Chiayi	Taiwan	Collapsed on July 1, 1997, following heavy rainstorms.[40]
Patriarca da Floresta [pt]	3,020	Jequitibá-rosa Cariniana legalis	Santa Rita do Passa Quatro, São Paulo	Brazil	Alive. Probably the oldest non-conifer in Brazil. Its name translates as "Patriarch of the Forest". Located at the Vassununga State Park.[41]
Great Camphor of Takeo	3,000+	Cinnamomum camphora "Camphor Tree"	Takeo, Saga	Japan	A large Camphor tree reputed to be over 3,000 years old.[42] Worshipped at a Shinto shrine dedicated to the tree, dating back to the 700s C.E.[43] Alive.
Raintree	3,000+	Great Basin bristlecone pine	Spring Mountains, Nevada	United States	Great Basin Bristlecone Pine located near Kyle Canyon in the Spring Mountain range in Southern Nevada, USA. Estimated 3000 years old but never cored.
Oliveira de Santa Iria de Azóia [Wikidata]	2,850	Olive Olea europaea	Santa Iria de Azóia [pt], Loures, Lisbon	Portugal	Magnificent Olive tree, probably the last one from a large olive grove. Studied by UTAD University and now classified "Public interest tree" by the Portuguese National Forest Authority; Tree ID[44]
Mother of the Forest	2,520	Giant sequoia Sequoiadendron giganteum	Sierra Nevada, Calaveras Big Trees State Park, California	United States	Dead.
Sarv Zibad	2,500	Mediterranean cypress Cupressus sempervirens	Sarv Zibad, Khorasan	Iran	Also called "holly Sarv".[28][29]
General Sherman	2,300–2,700	Giant sequoia Sequoiadendron giganteum	Giant Forest, Sequoia National Park, California	United States	Alive with a height of 83.8 meters (275 ft), a diameter of 11 m (36 ft) at its base, and an estimated bole volume of 1,487 m^3 (52,513 cu ft), it is the largest known living single-stem tree, and among the tallest, widest, and longest-lived of all trees on the planet.[45][46]
Kayano Ōsugi	2,300	Japanese cedar Cryptomeria japonica	Yamanaka Onsen, Ishikawa	Japan	Girth of 9.6 m. One of the four trees believed to be sacred in the precincts in a Shinto shrine. Its name translates as "Great Sugi of Kayano".
Oliveira de Santa Luzia	2,210	Olive Olea europaea	Santa Luzia, Tavira, Algarve	Portugal	Alive.[47]
Jōmon Sugi	2,170–7,200	Japanese cedar Cryptomeria japonica	Yakushima	Japan	Girth of 16.4 m. Exact dating is made difficult by the rotten core of the trunk. Its name is a reference to the Jōmon period of Japanese prehistory.[48][49][50]
Ballyconnell Yew	2,000–5,000	Yew Taxus baccata	Ballyconnell, Annagh	Ireland	Oldest tree in Ireland.^[citation needed]
Elia Vouvon	2,000–5,000	Olive Olea europaea	Kolymvari, Crete	Greece	Its name translates as "Olive Tree of Vouves".[51]
Castagnu dê Centu Cavaddi	2,000–4,000	Sweet chestnut Castanea sativa	Sicily	Italy	Its name translates as the "Hundred Horse Chestnut".
Ulleungdo Hyangnamu	2,000–3,000	Chinese juniper Juniperus chinensis	Ulleung-gun, Gyeongbuk	South Korea	Girth of 4.5 m. One of the main branches was broken in 1985 by typhoon "Brenda".
Stara Maslina	2,249 (2,016-2482)[52]	Olive Olea europaea	Stari Bar, Bar	Montenegro
Tnjri	2,040+	Oriental plane Platanus orientalis	Martuni Region	Azerbaijan	Oldest tree in a part of Azerbaijan. Its trunk is hollow. Also known as "Sose's Tree".
Koca Katran	2,030	Lebanon cedar Cedrus libani	Antalya	Turkey	Alive.[53]
?	2,010	Olive Olea europaea	Exo Hora, Zakynthos	Greece	[54]
?	2,010	Yew Taxus	Sochi, Krasnodar Krai	Russia	A grove known to possess several 2,000-year-old specimens.[55]
Houkisugi at Nakagawa	2,000	Japanese cedar Cryptomeria japonica	Nakagawa Settlement, Yamakita, Kanagawa	Japan
Lady Liberty	2,000+	Bald cypress Taxodium distichum	Longwood, Florida	United States	Sometimes called the "Companion Tree" to 'Senator".
Tejo Milenario	2,000+	Yew Taxus baccata	Cazorla, Jaén	Spain	Oldest tree in Spain.[56]
The Pechanga Great Oak Tree	1,000[57]-2,000	Coast live oak Quercus agrifolia	Temecula, California	United States	Oldest oak tree in the United States, possibly in the world.
Methuselah	1,800	Coast redwood Sequoia sempervirens	Woodside, California	United States	[58]
Araucaria Madre	1,810	Araucaria araucana	Araucanía	Chile	Located in Parque Nacional Conguillío. 1,800 rings account for at least 1,800 years.[59]
Granit oak	1,679	Pedunculate oak Quercus robur	Granit	Bulgaria
Kongeegen	1,500–2,000	Pedunculate oak Quercus robur	Jægerspris Nordskov, Zealand	Denmark	[60]
Stelmužė Oak	1,500–2,000	Pedunculate oak Quercus robur	Stelmužė, Zarasai	Lithuania	Oldest tree in the Baltic states. Supposedly oldest oak in Europe.
Vilémovice Yew	1,500–2,000	Common yew Taxus baccata	Vilémovice	Czech Republic	345 cm (136 in) in circumference.[61]
Aubépines	1,500	Common hawthorn Crataegus monogyna	Saint-Mars-sur-la-Futaie, Mayenne	France	Oldest tree in France.[62]
Jardine Juniper	1,500	Rocky Mountain juniper Juniperus scopulorum	Logan Canyon, Utah	United States	[63]
Árbol del Tule	1,433–1,600	Montezuma cypress Taxodium mucronatum	Santa María del Tule, Oaxaca	Mexico	Stoutest tree trunk in the world. Its name translates as the "Tule Tree".
Baikushev's pine	1,300	Heldreich's pine Pinus heldreichii	Pirin Mountains	Bulgaria	Oldest known coniferous tree in Bulgaria. It is named after its discoverer, forest ranger Kostadin Baikushev.
Te Matua Ngahere	1,200–4,000	Kauri Agathis australis	Waipoua Forest, Northland	New Zealand	Oldest tree in New Zealand. Its name translates as "Father of the Forest".
Cis Henrykowski	1270	Common Yew Taxus Baccata	Henryków Lubański, Lower Silesia,	Poland	Oldest tree in Poland. The age of the tree was estimated based on the trunk circumference and growth rate assessed by taking a core from an undamaged part of the trunk. If the trunk is the result of the fusion of two or three trunks, its age may be younger.[64]
Algarrobo Abuelo	1,200+	Algarrobo Prosopis chilensis	Villa de Merlo, San Luis Province	Argentina	Possibly oldest algarrobo in Argentina.^[citation needed]
"The Sisters" or "Sisters Olive Trees of Noah"	1,162 ±131[65]	Olive Olea europaea	Bchaaleh, North Governorate	Lebanon	Folk legend also ascribes The Sisters as the source of the olive branch returned to Noah's Ark at the waning of the Biblical Flood.[66]

Clonal trees

As with all long-lived plant and fungal species, no individual part of a clonal colony is alive (in the sense of active metabolism) for more than a very small fraction of the life of the entire clone. Some clonal colonies may be fully connected via their root systems, while most are not actually interconnected, but are genetically identical clones which populated an area through vegetative reproduction. Ages for clonal colonies, often based on current growth rates, are estimates.^[citation needed]

Name	Age (years)	Species	Location	Country	Notes
Antarctic Beech	2000	Antarctic beech Nothofagus moorei	Repeater Station Road, Springbrook National Park, Queensland	Australia	Three individuals approximately 2000 years old are the parent trees to others in the surrounding forest; they grow clones by coppicing new shoots radially, forming a ring at the base of the parent tree trunk.[67]
Wollemi Pines	60+ million years	Wollemi pine Wollemia nobilis	Wollemi National Park, New South Wales	Australia	According to Cris Brack and Matthew Brookhouse at the ANU Fenner School of Environment & Society: "Once you accept that a common, genetically identical stock can define a tree, then the absolute "winner" for oldest tree (or the oldest clonal material belonging to a tree) [in Australia] must go to the Wollemi pine (Wollemia nobilis). It may be more than 60 million years old. The Wollemi pine clones itself, forming exact genetic copies. It was thought to be extinct until a tiny remnant population was discovered in Wollemi National Park in 1994... There is also substantial evidence that the tree has been cloning itself and its unique genes ever since it disappeared from the fossil record more than 60 million years ago."[68]
Unknown	43,600–130,000	King's lomatia Lomatia tasmanica	Southwest National Park, Tasmania	Australia	Lomatia tasmanica is an endangered species with only one population, consisting of several hundred genetically identical individuals. Although it flowers occasionally, fruit production has never been observed, and it propagates vegetatively. Fossilized leaves are dated as at least 43,600 years old, found in sediments with a maximum age of 130,000.[69]
Pando	less than 14,000[4]–80,000[70]	Quaking aspen Populus tremuloides	Fishlake National Forest, Utah	United States	Covers 107 acres (0.43 km^2) and has around 47,000 stems (aged up to 130 years), which continually die and are renewed by its roots. Is also the heaviest-known organism, weighing 6,000 tonnes.
Jurupa Oak[71]	13,000[72]	Palmer oak Quercus palmeri	Jurupa Mountains, California	United States	Quercus palmeri Engelm. = Quercus dunnii Kellogg.[72]
Old Tjikko	9,550	Norway spruce Picea abies	Fulufjället National Park, Dalarna	Sweden	The tree's stems live no more than 600 years, but its root system's age[73][74] was established using carbon dating and genetic matching.[75] Elsewhere in the Fulu mountains, 20 spruces have been found older than 8,000 years.[76]
Old Rasmus	9,500	Norway spruce Picea abies	Sonfjället, Härjedalen	Sweden	[77]
Mongarlowe mallee	3,000–13,000	Mongarlowe mallee Eucalyptus recurva	Mongarlowe, New South Wales	Australia	Alive. A critically endangered species, known only from 6 wild individuals, first described in the 1980s. Based on the size and growth rate of its lignotuber, the largest single specimen is estimated to be 3,000 years old. However, it is possible that two other specimens are actually the result of a split in the original rootstock, and based on their spread of 26m would be estimated at 13,000 years old, potentially the oldest single tree on earth.[78]
unknown	3,000[79]–10,000[80]	Huon pine Lagarostrobos franklinii	Mount Read, Tasmania	Australia	Several genetically identical males that have reproduced vegetatively. Although single trees in this stand may be around 3 to 4 thousand years old, the stand itself as a single organism has existed for 10,000 years.[8]

See also

• King Clone
• List of largest giant sequoias
• List of longest-living organisms
• List of old-growth forests
• List of individual trees
• List of superlative trees
• Oldest viable seed
• Veteran tree

Notes

1. It is uncertain when Methuselah's age was determined, but "it seems likely the tree had 4,789 rings (crossdated) in the summer of 1957".[10] The age given here is based on this reference point.
2. Not named yet

References

1. https://www.biorxiv.org/content/10.1101/2024.10.19.619233v2.full.pdf
2. Farmer, Jared (2022). Elderflora: A Modern History of Ancient Trees. Basic Books. ISBN 978-0-465-09785-2.
3. Rogers, Paul C.; McAvoy, Darren J. (2018-10-17). "Mule deer impede Pando's recovery: Implications for aspen resilience from a single-genotype forest". PLOS ONE. 13 (10): e0203619. Bibcode:2018PLoSO..1303619R. doi:10.1371/journal.pone.0203619. ISSN 1932-6203. PMC 6192553. PMID 30332420.
4. Ding, Chen; Schreiber, Stefan G.; Roberts, David R.; Hamann, Andreas; Brouard, Jean S. (2017-07-05). "Post-glacial biogeography of trembling aspen inferred from habitat models and genetic variance in quantitative traits". Scientific Reports. 7 (1): 4672. Bibcode:2017NatSR...7.4672D. doi:10.1038/s41598-017-04871-7. ISSN 2045-2322. PMC 5498503. PMID 28680120.
5. Grant, Michael C. (1 October 1993). "The Trembling Giant". Discover Magazine. Archived from the original on 17 May 2012. Retrieved 8 May 2008.
6. "Botanical Record-Breakers". W.P Armstrong. Archived from the original on 19 December 2010. Retrieved 7 May 2008.
7. "SPECIES: Populus tremuloides". USDA - United States Forest Service. Archived from the original on 5 September 2012. Retrieved 8 May 2008.
8. "Native Conifers of Tasmania". Paks and Wildlife Service, Tasmania. Archived from the original on 28 July 2012. Retrieved 6 May 2008.
9. Salzer, Matthew; Baisan, Christopher (May 13–17, 2013). Dendrochronology of the "Currey Tree". Second American Dendrochronology Conference. University of Arizona, Tucson.
10. "Pinus longaeva". Gymnosperm Database. 2020-01-17. Retrieved 2021-10-17.
11. "OldList, A Database Of Old Trees". Rocky Mountain Tree-Ring Research. May 2019. Archived from the original on 12 April 2013. Retrieved 15 November 2019.
12. Lara, Antonio; Villalba, Ricardo (21 May 1993). "A 3620-Year Temperature Record from Fitzroya cupressoides Tree Rings in Southern South America". Science. 260 (5111): 1104–1106. Bibcode:1993Sci...260.1104L. doi:10.1126/science.260.5111.1104. PMID 17806339. S2CID 46397540.
13. "Is the world's oldest tree growing in a ravine in Chile?". Science Journal. 2022.
14. "Parque Nacional Alerce Costero". Conaf.cl (in Spanish). National Forest Corporation. Archived from the original on 6 October 2014. Retrieved 16 March 2014.
15. "Juniperus grandis". Gymnosperm Database. 2020-01-17. Retrieved 2021-10-17.
16. Stahle, D. W.; Edmondson, J. R.; Howard, I. M.; Robbins, C. R.; Griffin, R. D.; Carl, A.; Hall, C. B.; Stahle, D. K.; Torbenson, M. C. A. (May 16, 2019). "Longevity, climate sensitivity, and conservation status of wetland trees at Black River, North Carolina". Environmental Research Communications. 1 (4): 041002. Bibcode:2019ERCom...1d1002S. doi:10.1088/2515-7620/ab0c4a.
17. Brunstein, F. Craig; Yamaguchi, David K. (Aug 1992). "The Oldest Known Rocky Mountain Bristlecone Pines (Pinus aristata Engelm.)". Arctic and Alpine Research. 24 (3): 253–256. doi:10.2307/1551666. JSTOR 1551666. S2CID 128446269.
18. Adrian Patrut et al. (2018) The demise of the largest and oldest African baobabs. Nature Plants 4: 423–426. doi:10.1038/s41477-018-0170-5
19. "Oldest-known human-planted tree". Guinness world records. 2011.
20. Worrall, John (1990). "Subalpine Larch: Oldest Trees in Canada?". The Forestry Chronicle. 66 (5): 478–479. doi:10.5558/tfc66478-5.
21. Miles, D.H.; Worthington, M.J. (17–21 June 1998). "Sonora Pass junipers from California USA: construction of a 3,500-year chronology". In Stravinskiene, V.; Juknys, R. (eds.). Dendrochronology and Environmental Trends - Proceedings of the International Conference. Kaunas, Lithuania: Vytautas Magnas University Department of Environmental Sciences.
22. Suzuki, Eizi; Tsukahara, Jun (1987). "Age structure and regeneration of old growth Cryptomeria japonica forests on Yakushima Island". Journal of Plant Research. 100 (3): 223–241. doi:10.1007/BF02492832. S2CID 25942726.
23. Schulman, E. (1956). Dendroclimatic Changes in Semiarid America. Tucson: Univ. of Ariz. Press.
24. Larson, D. W.; Kelly, P. E. (1991). "The extent of old-growth Thuja occidentalis on cliffs of the Niagara Escarpment". Canadian Journal of Botany. 69 (7): 1628–1636. doi:10.1139/b91-206.
25. Stahle, D. W.; Cleaveland, M. K.; Hehr, J. G. (10 June 1988). "North Carolina Climate Changes Reconstructed from Tree Rings: A.D. 372 to 1985". Science. 240 (4858): 1517–1519. Bibcode:1988Sci...240.1517S. doi:10.1126/science.240.4858.1517. PMID 17798982. S2CID 20004819.
26. Piovesan, Gianluca; Biondi, Franco; Baliva, Michele; Saba, Emanuele Presutti; Calcagnile, Lucio; Quarta, Gianluca; D'Elia, Marisa; Vivo, Giuseppe De; Schettino, Aldo; Filippo, Alfredo Di (2018). "The oldest dated tree of Europe lives in the wild Pollino massif: Italus, a strip-bark Heldreich's pine". Ecology. 99 (7): 1682–1684. doi:10.1002/ecy.2231. ISSN 1939-9170. PMID 29768656.
27. "The Prometheus Story - Great Basin National Park (U.S. National Park Service)". www.nps.gov. Retrieved 2023-04-28. another bristlecone in the same area, proved to be 5,065 years old.
28. "Cypress". Encyclopædia Iranica. Archived from the original on 2017-11-17.
29. "Trees in ancient Iran". Earthwatchers. Archived from the original on 2013-01-17. Retrieved 2013-01-11.
30. "Oldest yew tree found in Turkey's north". 13 June 2016. Archived from the original on July 20, 2016. Retrieved July 9, 2016.
31. "Karadeniz Teknik Üniversitesi". www.ktu.edu.tr. Retrieved 2020-01-05.
32. "The ancient yew tree". St Johns Tisbury. Archived from the original on 10 September 2018. Retrieved 19 January 2019.
33. Pavuk, Amy. "'The Senator' Catches Fire". Orlando Sentinel. Times Newspaper. Archived from the original on 31 January 2012. Retrieved 16 January 2012.
34. Scheper, J. "Florida's Big Tree". Floridata.com. Archived from the original on 20 July 2008. Retrieved 7 May 2008.
35. "Archived copy" (PDF). Archived (PDF) from the original on 2016-11-04. Retrieved 2016-11-03.
36. "FLORESTAS — ICNF" (in Portuguese). Archived from the original on 2018-08-09. Retrieved 2016-11-03.
37. "A Luras il patriarca degli olivastri millenari d'Europa" (PDF) (in Italian). COMUNE DI LURAS. Archived from the original (PDF) on 2013-01-11.
38. Bevan-Jones, Robert (2004). The ancient yew: a history of Taxus baccata. Bollington: Windgather Press. pp. 38–39. ISBN 978-0-9545575-3-9.
39. Hidson, Toby (2014). "Addressing the claim that the Defynnog yewsin Powysmay be 5,000 years old" (PDF). Ancient Yew Group. Archived (PDF) from the original on December 24, 2018. Retrieved July 30, 2019.
40. "Alishan Forest Recreation area has new "tree king"". Taipei Times. 2006-07-02. Archived from the original on 2018-10-17. Retrieved 2013-01-11.
41. Botosso, Paulo Cesar (November 2002). "Conhecer a Idade das Árvores: Importância e Aplicação" (PDF). Embrapa Florestas. Documentos 75: 25. Archived (PDF) from the original on 2017-10-26. Retrieved 2017-10-26.
42. "A town with sacred camphor trees and a new-style library". Retrieved 2022-09-15.
43. "武雄神社とは". Retrieved 2022-09-15.
44. "UTAD classificou oliveira com 2.850 anos!" (in Portuguese). CienciaPT. 2011-07-08. Archived from the original on 2012-03-19. Retrieved 2013-01-11.
45. "The General Sherman Tree". Sequoia National Park. U.S. National Park Service. 1997-03-27. Retrieved 2011-08-12.
46. "The General Sherman Tree - Sequoia & Kings Canyon National Parks (U.S. National Park Service)". www.nps.gov. Archived from the original on 2018-03-17. Retrieved 2018-03-03.
47. "Árvores Monumentais de Portugal". ICNF. Retrieved 1 November 2020.
48. Hobson, Jake (2007). Niwaki: Pruning, Training and Shaping Trees the Japanese Way. Timber Press. p. 86. ISBN 978-0-88192-835-8. Retrieved 25 August 2008.
49. Flora and Fauna Fact Sheet (Web Japan)
50. Miyazaki, Makoto. "The old men of the mountains". The Yomiuri Shimbun. Archived from the original on 16 October 2007. Retrieved 15 June 2008.
51. "The Olive Tree and Olive Oil". Sfakia-crete.com. Archived from the original on 2018-12-12. Retrieved 2013-01-11.
52. "Can Montenegro's 2,200-Year-Old Olive Tree Be Saved?". 22 January 2024. Retrieved 2024-09-22.
53. "Locals want quarries out of Çığlıkara Forest". Retrieved July 9, 2016.
54. "the-oldest-olive-tree pictures, videos and albums". Webshots.com. Archived from the original on 2011-09-27. Retrieved 2013-01-11.
55. "О Кавказе". Wwf.ru. 2008-07-24. Archived from the original on 2016-12-16. Retrieved 2013-01-11.
56. "Andalucía alberga el árbol más longevo de toda España". Cambio16. 2018-05-04. Archived from the original on 2019-10-14. Retrieved 2019-10-15.
57. "Pechanga Band of Indians: The Great Oak". Retrieved 2024-09-22.
58. ""Methuselah" Redwood - Woodside, California". Waymarking.com. Archived from the original on 2018-08-09. Retrieved 2013-01-11.
59. ""Parque de Araucarias Araucanas" - Chile: La edad milenaria e indefinida de las Araucarias Araucanas". parquedearaucarias.blogspot.cl. 11 August 2013. Archived from the original on 2016-01-08. Retrieved 2015-12-30.
60. "Kong Frederik den Syvendes Stiftelse Paa Jægerspris". Kongfrederik.dk. Archived from the original on 2018-10-21. Retrieved 2013-01-11.
61. "Tajemný tis ve Vilémovicích je nejstarším stromem v Česku" (in Czech). Czech Radio. 2021-04-24. Retrieved 2023-06-20.
62. "Common Hawthorn next to the church in Saint-Mars-sur-la-Futaie, Mayenne, France".
63. "Jardine Juniper Trail". Utahmountainbiking.com. 2001-08-31. Archived from the original on 2019-03-10. Retrieved 2013-01-11.
64. Zarzyński, Paweł; Tomusiak, Robert. "European yew (Taxus baccata L.) from Henrykow Lubański – some doubts about the real age of the oldest tree in Poland" (PDF). Rocznik Polskiego Towarzystwa Dendrologicznego. 62: 49–52 – via Polskie Towarzystwo Botaniczne.
65. Camarero, J. Julio; Touchan, Ramzi; Valeriano, Cristina; Bashour, Isam; Stephan, Jean (April 2024). "Dating the Noah trees to improve age estimates in centennial and millennial olive trees". Dendrochronologia. 84: 126181. Bibcode:2024Dendr..8426181C. doi:10.1016/j.dendro.2024.126181. hdl:10261/354303. Retrieved 20 September 2024.
66. "Epic Olive Trees - Olive Oil Times". www.oliveoiltimes.com. Retrieved 2021-07-01.
67. "National Trust - Antarctic Beech (Nothofagus moorei )". trusttrees.org.au. Retrieved 9 April 2023.
68. "Where the old things are: Australia's most ancient trees". The Conversation. 18 April 2017. Retrieved 4 May 2023.
69. Lynch, A. J. J.; Barnes, R. W.; Vaillancourt, R. E.; Cambecèdes, J. (1998). "Genetic evidence that Lomatia tasmanica (Proteaceae) is an ancient clone" (PDF). Australian Journal of Botany. 46 (1): 25–33. doi:10.1071/BT96120. Retrieved 11 November 2013.
70. "Quaking Aspen". Bryce Canyon. National Park Service. Archived from the original on 2015-02-10.
71. Connor, Steve (December 23, 2009). "At 13,000 years, tree is world's oldest organism". The Independent. United Kingdom. Archived from the original on 2017-10-22.
72. May, Michael R; Provance, Mitchell C; Sanders, Andrew C; Ellstrand, Norman C; Ross-Ibarra, Jeffrey (December 2009). "A Pleistocene Clone of Palmer's Oak Persisting in Southern California". PLOS ONE. 4 (12): e8346. Bibcode:2009PLoSO...4.8346M. doi:10.1371/journal.pone.0008346. PMC 2796394. PMID 20041136.
73. "Swedes find 'world's oldest tree'". BBC News. 17 April 2008. Archived from the original on 21 April 2008. Retrieved 20 April 2008.
74. Owen, James (14 April 2008). "Oldest Living Tree Found in Sweden". National Geographic. Archived from the original on 18 April 2008. Retrieved 16 April 2008.
75. "World's oldest living tree discovered in Sweden". Swedish Research Council. 16 April 2008. Archived from the original on 3 October 2011. Retrieved 16 April 2008.
76. Roger Highfield (17 April 2008). "World's oldest tree discovered in Sweden". London: The Daily Telegraph. Archived from the original on 26 August 2008. Retrieved 15 December 2019.
77. "9 500 år gammal gran i Härjedalen". dn.se (in Swedish). Archived from the original on 2009-10-23.
78. "Eucalyptus recurva (a mallee) Draft Recovery Plan" (PDF). Sydney, Australia: National Parks and Wildlife Service NSW. 2002.
79. "The Oldest Living Tasmanian: The Huon Pine". Australian Broadcasting Company. Archived from the original on 2012-11-09. Retrieved 2013-01-11.
80. "Oldest Living Tree Found in Sweden". National Geographic News. 2008-04-08. Archived from the original on 2012-11-01. Retrieved 2013-01-11.

72. National Trust. 2023. Antarctic Beech (Nothofagus moorei). [ONLINE] Available at: https://trusttrees.org.au/tree/QLD/Springbrook/Repeater_Station_Road. [Accessed 8 April 2023].