Agua de la Piedra Formation

Agua de la Piedra Formation
Stratigraphic range: Late Oligocene (Deseadan) ~26–23 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Type	Geological formation
Unit of	Malargüe Group
Sub-units	"Rodados Lustrosos" level
Underlies	alluvium
Overlies	Pircala-Coihueco Formation
Thickness	37 metres (121 ft) (tuffs)
Lithology
Primary	Tuff
Other	Paleosols
Location
Coordinates	36.6°S 69.7°W / -36.6; -69.7
Approximate paleocoordinates	37.8°S 62.9°W / -37.8; -62.9
Region	southern Mendoza Province
Country	Argentina
Extent	southernmost Precordillera northernmost Neuquén Basin
Type section
Named by	Gorroño et al.
Location	Quebrada Fiera, Malargüe
Year defined	1979
Coordinates	36°33′13.3″S 69°42′3.5″W
Region	Mendoza Province
Country	Argentina
Thickness at type section	37 metres (121 ft) (tuffs)
Agua de la Piedra Formation (Argentina)

The Agua de la Piedra Formation (FAP, Spanish names include Estratos de Agua de la Piedra and Complejo Volcano-sedimentario del Terciario inferior)[1] is a Late Oligocene (Deseadan in the SALMA classification) geologic formation of the Malargüe Group that crops out in the southernmost Precordillera and northernmost Neuquén Basin in southern Mendoza Province, Argentina.[2]

The strictly terrestrial tuffs and paleosols of the formation, geologically belonging to Patagonia, have provided a wealth of mammal fossils of various groups at Quebrada Fiera, including Mendozahippus fierensis, Pyrotherium, Coniopternium and Fieratherium. Terror birds reminiscent of the terror bird Andrewsornis and indeterminate remains of the phorusrhacid family have found in conjunction with the mammals.

Regional geology

The Agua de la Piedra is geologically part of the Neuquén Basin, Argentina's most prolific onshore petroleum producing basin of northwestern Patagonia, and crops out in the geographical feature of the Andean orogeny; the Argentinian Precordillera of the higher Andes in the hinterland. The Malargüe Group, of which the Agua de la Piedra Formation is the uppermost unit, hosts among the most spectacular dinosaur fossils and nesting sites in the Allen Formation, the lowermost stratigraphic unit of the group.

The Jagüel Formation, overlying the Allen Formation, hosts the Cretaceous–Paleogene boundary and has provided fossils of marine reptiles including mosasaurs[3] and the marine turtle Euclastes meridionalis. The Roca Formation, overlying the Jagüel Formation shows evidence of Atlantic waters depositing the evaporites, claystones and limestones of the formation.[4][5]

The Neuquén Basin started forming in the latest Jurassic as one of the rift basins resulting from the break-up of Pangea. While the earlier formations in the basin are mostly distal terrestrial in nature, the Agua de la Piedra Formation is a unique combination of purely terrestrial influence (paleosols) with the early Andean volcanism in the form of tuffs.

Oligocene South America

See also: Category:Oligocene Series of South America and Category:Paleogene Argentina

Climate

Global cooling occurred during the Oligocene

Eocene-Oligocene circum-Antarctic oceanic changes

Oligocene South America differed quite substantially from the Eocene period preceding it. Isolated from Gondwana for 70 million years, the continent had developed widespread lush forests with their own specific faunas. The climate drastically cooled at the Eocene-Oligocene boundary with global cooling as a result of the formation of the Antarctic Ocean current. The South American landscape became more arid than in the Eocene with ongoing volcanism related to the Andean orogeny affecting the local climates.

Oligocene fauna

The Oligocene of South America is characterized by the arrival of the first monkeys, possibly rafting from Africa, which in the Oligocene was significantly removed from South America. The first rodents had arrived to the island continent in the Late Eocene before,[6] perhaps using similar methods of transoceanic transport. The rodents of South America diversified in the Oligocene. Cabeza Blanca, where the Sarmiento Formation outcrops, has provided the richest and most diverse Oligocene fauna of South America.[7]

The cooler Oligocene climate led to the widespread extension of savanna and other grassland biomes. In the Early Oligocene, these rodents inhabited open and arid landscapes with wind-blown dust and grasslands environments.[8]

Monkeys and rodents

The oldest confirmed New World monkey fossils stem from the Deseadan formations Salla in presently Andean Bolivia (the approximately 1,000 grams (2.2 lb) weighing Branisella boliviana and Szalatavus attricuspis half the size of Branisella) and the 2,000 g (4.4 lb) heavy Canaanimico from the Chambira of Amazonian Peru.[9]

The rodents had arrived in the Late Eocene and diversified greatly during the Deseadan following the appearance of Andemys with species A. frassinettii and A. termasi in the Tinguirirican (Abanico Formation; Tinguiririca fauna). Caviomorphs arrived in Patagonia during the latest Eocene or early Oligocene, and by the Late Oligocene they were highly diversified, with representatives of the four main lineages. A great morphological disparity, at least in tooth morphology, was then acquired mainly by the development of hypsodonty in several lineages. The early evolution of each of the major clades was complex, especially for chinchilloids and octodontoids. The first stages of the evolution of cavioids are more obscure because they are recognized through the relatively derived Deseadan species of Cavioidea.[10]

The Oligocene (Tinguirirican and Deseadan SALMAs plus La Cantera fauna) has a rich record of caviomorphs showing a greater morphological disparity than older faunas. Representatives of the four superfamilies, with the archetypal dental features that characterize species of the subsequent SALMAs, can be clearly recognized, at least since the Deseadan SALMA. Although a few genera (e.g., Andemys, Branisamys) cannot be assigned with certainty to any supra generic taxa. The Acaremyidae were likely a group of austral differentiation. The first representatives, the Deseadan Platypittamys brachyodon, Galileomys baios and Changquin woodi,[11] attest to its differentiation into several lineages.[12]

Oligocene volcanism

Payún Matrú volcano

Early Andean volcanism in the Southern Cone of South America dating to the Oligocene has been found in:

• Estratos de Pupunahue[13][14][15]
• Estratos de San Pedro[13]
• Caleta Chonos Formation
• Cheuquemó Formation
• Temuco Formation[16]
• Oxaya Formation[17][18]
• Ancud Volcanic Complex
• Cerro Ratones

Description

The formation comprises the "Rodados Lustrosos" level, formed by clastic heterogeneous conglomerates in a silty matrix, considered as the stratigraphic evidence of the Pehuenche orogenic phase of the Andean orogeny, followed by uniform sequences, variable in thickness, of whitish-ocher tuffaceous paleosols with concretions and whitish-gray tuffs with intercalations of pyroclastic deposits.[19]

The upper part of the Agua de la Piedra Formation consists of 37 metres (121 ft) of white-grayish tuffs and tobaceous paleosols, with laminated or massive parallel stratification constitute the fossiliferous level of Quebrada Fiera.[20] The formation overlies the Pircala-Coihueco Formation.[21]

Depositional environment

The studied profiles of the Agua de la Piedra Formation show large lateral lithological varieties, typical of alluvial fan depositional setings. The climate during deposition has been estimated to be semi-arid and the differential thicknesses of facies associations within the Agua de la Piedra Formation may represent the infill of minibasins in the forming foreland of the Andes. Sedimentary loading can enhance the effect of tectonic forces in foreland basins. The variety in volcanic fragments and composition indicates local ash fall caused by contemporaneous volcanism in the area of deposition.[22]

2017 research on the Deseadan fauna (late Oligocene) from Quebrada Fiera, south of Mendoza Province, Argentina, evidences a rich mammal assemblage that shows the existence of common elements with Deseadan faunal associations of Patagonia and those of lower latitudes such as Salla, Bolivia, as well as endemic taxa of different groups.[23]

Endemism refers to Notohippidae (Mendozahippus fierensis), Leontiniidae (Gualta cuyana), Homalodotheriidae (Asmodeus petrasnerus) and Metatheria (Fieratherium sorex); to these mammals a new terrestrial snail has been added in 2016.[24]

Faunal data published in 2019 confirm the Deseadan age, but as per 2020, absolute dating is lacking for Quebrada Fiera.[19]

Paleontological significance

Quebrada Fiera

The Quebrada Fiera site is situated in the Malargüe Department,[25] southern Mendoza Province, Argentina, in the foothills of the Andes Range. The fossiliferous levels are located at around 36°33′13.3″S 69°42′3.5″W at 1,300 to 1,406 metres (4,265 to 4,613 ft) elevation. The site was discovered during a geological prospection carried out by Yacimientos Petrolíferos Fiscales (YPF) in the late 1970s (Gorroño et al., 1979). Later on, other fossil bearing levels were found at the southern side of the ravine,[26] located at around 36°33′26″S 69°41′35″W, 1,316 metres (4,318 ft) elevation.[19]

The site is one of five recognized fossiliferous sites in Mendoza Province, with Divisadero Largo, where the Santacrucian Mariño Formarion is found, Huaquerías, defining the Huayquerian in the Huayquerías Formation, the Aisol Formation of central Mendoza and the Uspallata Group and Carrizal Formations in the north of the province.[21]

The geological characterization and the preliminary faunal list were published by Gorroño et al. (1979). The faunal assemblage was then assigned to the Late Oligocene (Deseadan SALMA) based on the presence of two typical representatives of the Deseadean fauna of Patagonia; Pyrotherium Ameghino 1888 and Proborhyaena gigantea Ameghino 1897,[19] both also found in the Puesto Almendra member of the Sarmiento Formation.[27]

The species epithet Mendozahippus fierensis and genus Fieratherium refer to Quebrada Fiera.[25][26][28][29][30]

Fossil content

The formation has provided fossils of:[2]

Group	Clade	Taxa	Site	Images	Notes
Ungulates	Macraucheniidae	Coniopternium andinum	Quebrada Fiera North		[25][31]
	Proterotheriidae	cf. Lambdaconus suinus	Quebrada Fiera North		[25][32]
	Pyrotheriidae	Pyrotherium romeroi	Quebrada Fiera North		[25][33]
		Pyrotherium sp.	Quebrada Fiera South		[26]
	Litopterna	Litopterna indet.	Quebrada Fiera North		[25]
Cingulata	Dasypodidae	Meteutatus aff. lagenaformis	Quebrada Fiera North		[25][34]
		?Prozaedyus aff. impressus	Quebrada Fiera North		[25][34]
		Stenotatus aff. ornatus	Quebrada Fiera North		[25][34]
Xenarthra	Glyptodontinae	Glyptodontinae indet.	Quebrada Fiera North		[25]
	Megalonychidae	?Megalonychidae indet.	Quebrada Fiera North		[25]
Notoungulata	Notohippidae	Mendozahippus fierensis	Quebrada Fiera South		[26]
			Quebrada Fiera North		[25][35]
		Notohippidae indet.	Quebrada Fiera North		[25][36]
	Archaeohyracidae	cf. Archaeotypotherium sp.	Quebrada Fiera North		[25][37]
		Archaeohyrax suniensis	Quebrada Fiera North		[25][37][38]
	Hegetotheriidae	Prosotherium garzoni	Quebrada Fiera North		[25][39]
		cf. Prosotherium sp.	Quebrada Fiera North		[25]
		Prohegetotherium malalhuense	Quebrada Fiera North		[25]
		P. schiaffinoi	Quebrada Fiera North		[25]
		P. cf. sculptum	Quebrada Fiera North		[25]
		Prohegetotherium sp.	Quebrada Fiera North		[25]
		Hegetotheriopsis sulcatus	Quebrada Fiera North		[25][40]
	Homalodotheriidae	Asmodeus petrasnerus	Quebrada Fiera North		[25][29][41]
	Interatheriidae	Argyrohyrax proavus	Quebrada Fiera North		[25]
		Progaleopithecus sp.	Quebrada Fiera North		[25]
		Interatheriidae indet.	Quebrada Fiera South		[26]
	Leontiniidae	Gualta cuyana	Quebrada Fiera North		[25][42]
	Mesotheriidae	Trachytherus cf. spegazzinianus	Quebrada Fiera North		[25][43]
	Toxodontidae	Proadinotherium sp.	Quebrada Fiera North		[25][44]
		Toxodontidae indet.	Quebrada Fiera North		[25][45]
Rodents	Acaremyidae	Acaremyidae indet.	Quebrada Fiera North		[25]
Sparassodonta	Borhyaenidae	Pharsophorus sp.	Quebrada Fiera North		[25]
	Proborhyaenidae	Proborhyaena gigantea	Quebrada Fiera North		[25]
Theriiformes		Fieratherium sorex	Quebrada Fiera North		[25][46]
Birds	Phorusrhacidae	cf. Andrewsornis sp.	Quebrada Fiera North		[25]
		Phorusrhacidae indet.	Quebrada Fiera South		[26]
Invertebrates	Gastropods	Gastropoda indet.	Quebrada Fiera North		[25]

SALMA correlations

Main articles: Deseadan and South American land mammal age

The Deseadan South American land mammal age (SALMA) is equivalent to the Arikareean in the North American land mammal age (NALMA) and the Harrisonian in the 2000 version of the classification. It overlaps with the Hsandagolian of Asia and the MP 25 zone of Europe, the Waitakian and the Landon epoch of New Zealand.

Deseadan correlations in South America

Formation		Rancahué	Guillermo	Mariño	Deseado	Sarmiento	Salla	Lacayani	Fray Bentos	Moquegua	Chambira	Barzalosa	Tremembé	Cascadas	Map
Basin	Neuquén		Austral	Cuyo	Deseado	San Jorge	Salla	Subandean	Norte	Moquegua	Ucayali	VSM	Taubaté	Panama	Agua de la Piedra Formation (South America)
Country	Argentina						Bolivia		Uruguay	Peru		Colombia	Brazil	Panama
Archaeohyrax
Prohegetotherium
Pyrotherium
Pharsophorus
Trachytherus
Proadinotherium
Proborhyaena
Meteutatus
Andrewsornis
Terror birds
Rodents
Reptiles
Primates
Flora
Insects
Environments	Alluvial		Fluvial	Eolian Alluvial-fluvial	Fluvial		Alluvial	Fluvial-alluvial	Fluvial	Fluvio-lacustrine		Alluvial-fluvial	Lacustrine	Fluvial	Deseadan volcanoclastics Deseadan fauna Deseadan flora
Volcanic	Yes					Yes	Yes	Yes		Yes				Yes

See also

• Abanico Formation
• Dos Bocas Formation
• Collón Curá Formation
• Gaiman Formation
• Honda Group
• Itaboraí Formation
• Ituzaingó Formation
• Laguna del Hunco Formation
• Deseadan formations

References

1. Combina et al., 1994, p.418
2. Agua de la Piedra Formation in the Paleobiology Database
3. Hoja 3969-II Neuqúen, 2007
4. Archuby et al., 2016
5. Malamuián & Náñez, 2011
6. Vassallo & Antenucci, 2015, p.6
7. Vucetich et al., 2015, p.21
8. Ojeda et al., 2015, p.123
9. Silvestro et al., 2017, p.14
10. Vucetich et al., 2015, p.11
11. Vucetich et al., 2014, p.692
12. Vucetich et al., 2015, p.18
13. Elgueta et al., 2000
14. Alfaro & Gantz, 1997
15. Villablanca et al., 2003
16. Mella & Quiroz, 2010
17. García et al., 1999
18. Zeilinger et al., 2015
19. Schmidt et al., 2019, p.370
20. Cerdeño, 2012, p.378
21. Cerdeño, 2012, p.376
22. Combina et al., 1994, p.420
23. Hernández Pino et al., 2017, p.195
24. Miquel & Cerdeño, 2016
25. Quebrada Fiera at Fossilworks.org
26. Quebrada Fiera South in the Paleobiology Database
27. Gran Blanca in the Paleobiology Database
28. Cerdeño & Reguero, 2015
29. Seoane & Cerdeño, 2014
30. Cerdeño & Vera, 2014a
31. Schmidt et al., 2019, p.371
32. Schmidt et al., 2019, p.375
33. Cerdeño & Vera, 2017
34. Carlini et al., 2009
35. Cerdeño & Vera, 2010
36. Cerdeño & Vera, 2014b
37. Cerdeño et al., 2010
38. Vera et al., 2017
39. Seoane et al., 2019
40. Kramarz & Bond, 2017
41. Hernández Pino et al., 2017, p.206
42. Hernández Pino et al., 2017, p.201
43. Cerdeño, 2014
44. Hernández Pino et al., 2017, p.198
45. Hernández Pino et al., 2017, p.200
46. Forasiepi et al., 2014

Bibliography

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Regional geology

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Oligocene volcanism

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Paleontology

• Schmidt, Gabriela I.; Cerdeño, Esperanza; Hernández del Pino, Santiago (2019), "Macraucheniidae and Proterotheriidae (Mammalia, Litopterna) from Quebrada Fiera (Late Oligocene), Mendoza Province, Argentina", Andean Geology, 46 (2): 368–382, doi:10.5027/andgeoV46n2-3109, hdl:11336/80129, retrieved 2020-08-11 Material was copied from this source, which is available under a Creative Commons Attribution 3.0 International License.
• Seoane, F. D.; Cerdeño, E.; Singleton, H. (2019), "Re-assessment of the Oligocene genera Prosotherium and Propachyrucos (Hegetotheriidae, Notoungulata)", Comptes Rendus Palevol, 18 (6): 643–662, Bibcode:2019CRPal..18..643S, doi:10.1016/j.crpv.2019.07.001
• Cerdeño, Esperanza; Vera, Bárbara (2017), "New anatomical data on Pyrotherium (Pyrotheriidae) from the late Oligocene of Mendoza, Argentina", Ameghiniana, 54 (3): 290–306, doi:10.5710/AMGH.14.11.2016.3052, hdl:11336/63606
• Vera, B.; Cerdeño, E.; Reguero, M. (2017), "The Interatheriinae from the Late Oligocene of Mendoza (Argentina), with comments on some Deseadan Interatheriidae", Historical Biology, 29 (5): 607–626, Bibcode:2017HBio...29..607V, doi:10.1080/08912963.2016.1220945, hdl:11336/49742
• Hernández Del Pino, Santiago; Seoane, Federico D.; Cerdeño, Esperanza (2017), "New postcranial remains of large toxodontian notoungulates from the late Oligocene of Mendoza, Argentina and their systematic implications" (PDF), Acta Palaeontologica Polonica, 62: 195–210, retrieved 2019-02-12 Material was copied from this source, which is available under a Creative Commons Attribution 4.0 International License.
• Kramarz, A. G.; Bond, M. (2017), "Systematics and stratigraphical range of the hegetotheriids Hegetotheriopsis sulcatus and Prohegetotherium sculptum (Mammalia: Notoungulata)", Journal of Systematic Palaeontology, 15 (12): 1027–1036, Bibcode:2017JSPal..15.1027K, doi:10.1080/14772019.2016.1266047, hdl:11336/46835
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• Forasiepi, A. M.; Goin, F. J.; Abello, M. A.; Cerdeño, E. (2014), "A unique, Late Oligocene shrew-like marsupial from western Argentina and the evolution of dental morphology", Journal of Systematic Palaeontology, 12 (5): 549–564, Bibcode:2014JSPal..12..549F, doi:10.1080/14772019.2013.799611, hdl:11336/32498
• Seoane, Federico; Cerdeño, Esperanza (2014), "First extra-Patagonian record of Asmodeus Ameghino (Notoungulata, Homalodotheriidae) in the Late Oligocene of Mendoza Province, Argentina", Ameghiniana, 51 (5): 373–384, doi:10.5710/AMGH.07.07.2014.2735, retrieved 2019-02-12
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• Cerdeño, Esperanza (2012), "Quebrada Fiera (Mendoza), an important paleobiogeographic center in the South American late Oligocene", Estudios Geológicos, 67: 375–384, doi:10.3989/egeol.40519.194, retrieved 2020-08-11
• Cerdeño, Esperanza; Vera, Bárbara (2010), "Mendozahippus fierensis, gen. et sp. nov., new Notohippidae (Notoungulata) from the late Oligocene of Mendoza (Argentina)", Journal of Vertebrate Paleontology, 30 (6): 1805–1817, Bibcode:2010JVPal..30.1805C, doi:10.1080/02724634.2010.520781
• Cerdeño, Esperanza; Reguero, M.; Vera, B. (2010), "Deseadan Archaeohyracidae (Notoungulata) from Quebrada Fiera (Mendoza, Argentina) in the Paleobiogeographic Context of the South American Late Oligocene", Journal of Paleontology, 84 (6): 1177–1187, doi:10.1666/10-024.1
• Carlini, A.A.; Ciancio, M.R.; Flynn, J. J.; Scillato Yané, G. J.; Wyss, A. R. (2009), "The phylogenetic and biostratigraphic significance of new armadillos (Mammalia, Xenarthra, Dasypodidae, Euphractinae) from the Tinguirirican (early Oligocene) of Chile", Journal of Systematic Palaeontology, 7 (4): 489–503, Bibcode:2009JSPal...7..489C, doi:10.1017/S1477201908002708
• Patterson, Bryan; Marshall, Larry G. (1978), "The Deseadan, Early Oligocene, Marsupialia South America", Fieldiana: Geology, 41 (2): 37–100

New World monkeys

• Croft, Darin A (2016), Horned Armadillos and Rafting Monkeys: The Fascinating Fossil Mammals of South America, Indiana University Press, pp. 1–320, ISBN 9780253020949, retrieved 2017-10-21
• Fleagle, John G.; Rosenberger, Alfred L. (2013), The Platyrrhine Fossil Record, Elsevier, pp. 1–256, ISBN 9781483267074, retrieved 2017-10-21
• Hartwig, W.C.; Meldrum, D.J. (2002), The Primate Fossil Record - Miocene platyrrhines of the northern Neotropics, Cambridge University Press, pp. 175–188, ISBN 978-0-521-08141-2, retrieved 2017-09-24

Terror birds

• Sedor, Fernando A.; Vieira Dias, Eliseu; Da Cunha, Renata Floriano; Alvarenga, Herculano (2014), Paleogene phorusrhacid bird (Aves, Phorusrhacidae) from the Guabirotuba Formation, Curitiba Basin, Paraná, South of Brazil, 4th International Palaeontological Congress, p. 807, retrieved 2019-03-04
• Tambussi, C.P.; de Mendoza, R.; Degrange, F.J.; Picasso, M.B. (2013), "Flexibility along the Neck of the Neogene Terror Bird Andalgalornis steulleti (Aves Phorusrhacidae)", PLoS ONE, 7 (5): e37701, doi:10.1371/journal.pone.0037701, PMC 3360764, PMID 22662194
• Cenizo, Marcos M (2012), "Review Of The Putative Phorusrhacidae From The Cretaceous And Paleogene Of Antarctica: New Records Of Ratites And Pelagornithid Birds" (PDF), Polish Polar Research, 33 (3): 239–258, doi:10.2478/v10183-012-0014-3
• Alvarenga, H.; Jones, W.; Rinderknecht, A. (2010), "The youngest record of phorusrhacid birds (Aves, Phorusrhacidae) from the late Pleistocene of Uruguay" (PDF), Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen, 256 (2): 229–234, doi:10.1127/0077-7749/2010/0052
• Wroe, Stephen (2010), "Mechanical Analysis Of Feeding Behavior In The Extinct "Terror Bird' Andalgalornis steulleti (Gruiformes: Phorusrhacidae)", PLoS ONE, 5 (8): 1–7, Bibcode:2010PLoSO...511856D, doi:10.1371/journal.pone.0011856, PMC 2923598, PMID 20805872
• Alvarenga, Herculano M.F.; Höfling, Elizabeth (2003), "Systematic revision of the Phorusrhacidae (Aves: Ralliformes)", Papéis Avulsos de Zoologia, 43 (4): 55–91, doi:10.1590/S0031-10492003000400001

South American rodents

• Boivin, Myriam; Marivaux, Laurent; Orliac, Maëva J.; Pujos, François; Salas Gismondi, Rodolfo; Tejada Lara, Julia V.; Antoine, Pierre-Olivier (2017), "Late middle Eocene caviomorph rodents from Contamana, Peruvian Amazonia", Palaeontologia Electronica, 20 (1): Article number 20.1.19A, retrieved 2019-02-12
• Vassallo, Aldo I.; Antenucci, Daniel (2015), Biology of Caviomorph Rodents: Diversity and Evolution, SAREM, pp. 1–330, ISBN 9789879849736
  • Vassallo, Aldo I.; Antenucci, Daniel (2015), "Caviomorph rodents: an introduction", SAREM Series A, Prologue: 2–10
  • Vucetich, María G.; Arnal, Michelle; Deschamps, Cecilia M.; Pérez, Maria E.; Vieytes, Emma C. (2015), "A brief history of caviomorph rodents as told by the fossil record", SAREM Series A, Chapter 1: 11–63
  • Ojeda, Ricardo A.; Novillo, Agustina; Ojeda, Agustina A. (2015), "Large-scale richness patterns, biogeography and ecological diversification in caviomorph rodents", SAREM Series A, Chapter 3: 121–138
• Bertrand, Ornella C.; Flynn, John J.; Croft, Darin A.; Wyss, André R. (2012), "Two new taxa (Caviomorpha, Rodentia) from the early Oligocene Tinguiririca fauna (Chile)", American Museum Novitates (3750): 1–36, doi:10.1206/3750.2, retrieved 2019-02-15
• Antoine, P.O.; Marivaux, L.; Croft, D.A.; Billet, G.; Ganerod, M.; Jaramillo, C.; Martin, T.; Orliac, M.; Tejada, A. J. Altamirano, F. Duranthon, G. Fanjat, S. Rousse and R. Salas Gismondi, J. (2011), "Middle Eocene rodents from Peruvian Amazonia reveal the pattern and timing of caviomorph origins and biogeography", Proceedings of the Royal Society B, 279 (1732): 1319–1326, doi:10.1098/rspb.2011.1732, PMC 3282368, PMID 21993503, retrieved 2019-02-14
• Cerdeño, Esperanza; Guiomar Vucetich, M. (2007), "The first rodent from the Mariño Formation (Miocene) at Divisadero Largo (Mendoza, Argentina) and its biochronological implications" (PDF), Andean Geology, 34: 199–207, doi:10.5027/andgeoV34n2-a02, retrieved 2017-10-23

Regional correlations

Mariño Formation

• Cerdeño, Esperanza; Vera, Bárbara; Combina, Ana María (2018), "A new early Miocene Mesotheriidae (Notoungulata) from the Mariño Formation (Argentina): Taxonomic and biostratigraphic implications", Journal of South American Earth Sciences, 88: 118–131, Bibcode:2018JSAES..88..118C, doi:10.1016/j.jsames.2018.06.016, hdl:11336/158508, retrieved 2019-02-11

Rancahué Formation

• Vera, E.I (2010), "Oligocene ferns from the Rancahué Formation (Aluminé, Neuquén, Argentina): Cuyenopteris patagoniensis nov. gen., nov. sp. (Polypodiales: Blechnaceae/Dryopteridaceae) and Alsophilocaulis calveloi Menéndez emend. Vera (Cyatheales: Cyatheaceae)", Geobios, 43 (4): 465–478, Bibcode:2010Geobi..43..465V, doi:10.1016/j.geobios.2009.12.003
• Menéndez, C.A (1961), "Estípite petrificado de una nueva Cyatheaceae del Terciario de Neuquén", Boletín de la Sociedad Argentina de Botánica, IX: 331–358

Río Guillermo Formation

• Vento, B.; Gandolfo, M. A.; Nixon, K. C.; Prámparo, M. (2017), "Paleofloristic assemblage from the Paleogene Río Guillermo Formation, Argentina: preliminary results of phylogenetic relationships of Nothofagus in South America", Historical Biology, 29 (1): 93–107, Bibcode:2017HBio...29...93V, doi:10.1080/08912963.2015.1136930, hdl:11336/63607

Deseado Formation

• Agnolin, Federico (2006), "Posición sistemática de algunas aves fororracoideas (Ralliformes: Cariamae) Argentinas", Revista del Museo de Ciencias Naturales, 8: 27–33, doi:10.22179/REVMACN.8.348, retrieved 2018-09-10
• Forasiepi, Analía M.; Babot, M. Judith; Zimicz, Natalia (2014), "Australohyaena antiqua (Mammalia, Metatheria, Sparassodonta), a large predator from the Late Oligocene of Patagonia", Journal of Systematic Palaeontology, 13 (6): 503–525, Bibcode:2015JSPal..13..503F, doi:10.1080/14772019.2014.926403, hdl:11336/59430, retrieved 2019-02-12

Sarmiento Formation

• Acosta Hospitaleche, Carolina; Tambussi, Claudia (2005), "Phorusrhacidae Psilopterinae (Aves) en la Formación Sarmiento de la localidad de Gran Hondonada (Eoceno Superior), Patagonia, Argentina [Phorusrhacidae Psilopterinae (Birds) in the Sarmiento Formation from the Gran Hondonada locality (Upper Eocene), Patagonia, Argentina]" (PDF), Revista Española de Paleontología, 20: 127–132, retrieved 2018-09-10
• Arnal, Michelle; Kramarz, Alejandro G.; Vucetich, M. Guiomar; Vieytes, E. Carolina (2014), "A new early Miocene octodontoid rodent (Hystricognathi, Caviomorpha) from Patagonia (Argentina) and a reassessment of the early evolution of Octodontoidea", Journal of Vertebrate Paleontology, 34 (2): 397–406, Bibcode:2014JVPal..34..397A, doi:10.1080/02724634.2013.808203, hdl:11336/29748, retrieved 2019-02-12
• Cheme Arriaga, Lucas; Dozo, María Teresa; Gelfo, Javier N. (2016), "A new Cramaucheniinae (Litopterna, Macraucheniidae) from the early Miocene of Patagonia, Argentina", Journal of Vertebrate Paleontology, 36 (6): e1229672, Bibcode:2016JVPal..36E9672C, doi:10.1080/02724634.2017.1229672, hdl:11336/30386, retrieved 2019-02-12
• Dozo, María Teresa; Ciancio, Martín; Bouza, Pablo; Martínez, Gastón (2014), "Nueva asociación de mamíferos del Paleógeno en el este de la Patagonia (provincia de Chubut, Argentina): implicancias biocronológicas y paleobiogeográficas" (PDF), Andean Geology, 41: 224–247, doi:10.5027/andgeoV41n1-a09, retrieved 2017-08-15
• Pérez, María Encarnación; Krause, Marcelo; Vucetich, María Guiomar (2012), "A new species of Chubutomys (Rodentia, Hystricognathi) from the late Oligocene of Patagonia and its implications on the early evolutionary history of Cavioidea sensu stricto", Geobios, 45 (6): 573–580, Bibcode:2012Geobi..45..573P, doi:10.1016/j.geobios.2012.06.001
• Shockey, Bruce J.; Flynn, John J.; Croft, Darin A.; Gans, Phillip; Wyss, André R. (2012), "New leontiniid Notoungulata (Mammalia) from Chile and Argentina : comparative anatomy, character analysis, and phylogenetic hypotheses", American Museum Novitates (3737): 1–64, doi:10.1206/3737.2, retrieved 2019-02-15
• Sterli, Juliana; De la Fuente, Marcelo S.; Krause, J. Marcelo (2015), "A new turtle from the Palaeogene of Patagonia (Argentina) sheds new light on the diversity and evolution of the bizarre clade of horned turtles (Meiolaniidae, Testudinata)", Zoological Journal of the Linnean Society, 174 (3): 519–548, doi:10.1111/zoj.12252, hdl:11336/41594, retrieved 2019-02-13
• Vucetich, M.G.; Dozo, M.T.; Arnal, M.; Pérez, M.E. (2015), "New rodents (Mammalia) from the late Oligocene of Cabeza Blanca (Chubut) and the first rodent radiation in Patagonia", Historical Biology: An International Journal of Paleobiology, 27 (2): 236–257, Bibcode:2015HBio...27..236V, doi:10.1080/08912963.2014.883506, hdl:11336/17958, retrieved 2019-02-13
• Vucetich, María G.; Pérez, María E.; Ciancio, Martín R.; Carlini, Alfredo A.; Madden, Richard H.; Kohn, Matthew J. (2014), "A new acaremyid rodent (Caviomorpha, Octodontoidea) from Scarritt Pocket, Deseadan (late Oligocene) of Patagonia (Argentina)", Journal of Vertebrate Paleontology, 34 (3): 689–698, Bibcode:2014JVPal..34..689V, doi:10.1080/02724634.2013.804414, hdl:11336/17973, retrieved 2019-02-12
• Wyss, André R.; Flynn, John J.; Croft, Darin A. (2018), "New Paleogene notohippids and leontiniids (Toxodontia; Notoungulata; Mammalia) from the Early Oligocene Tinguiririca Fauna of the Andean Main Range, central Chile", Journal of South American Earth Sciences, 3903: 1–42, hdl:2246/6901, retrieved 2019-02-11

Salla Formation

• Reguero, Marcelo A.; Cerdeño, Esperanza (2005), "New late Oligocene Hegetotheriidae (Mammalia, Notoungulata) from Salla, Bolivia", Journal of Vertebrate Paleontology, 25 (3): 674–684, doi:10.1671/0272-4634(2005)025[0674:NLOHMN]2.0.CO;2, retrieved 2018-09-11
• Rincón, Ascanio D.; Shockey, Bruce J.; Anaya, Federico; Solórzano, Andrés (2015), "Palaeothentid Marsupials of the Salla Beds of Bolivia (Late Oligocene): Two New Species and Insights into the Post-Eocene Radiation of Palaeothentoids", Journal of Mammalian Evolution, 22 (4): 455–471, doi:10.1007/s10914-015-9295-8, retrieved 2019-02-13
• Shockey, Bruce J (2017), "New early diverging cingulate (Xenarthra: Peltephilidae) from the Late Oligocene of Bolivia and considerations regarding the origin of crown Xenarthra", Bulletin of the Peabody Museum of Natural History, 58 (2): 371–396, retrieved 2019-02-12

Lacayani fauna

• Billet, Guillaume; De Muizon, Christian; Mamani Quispe, Bernardino (2008), "Late Oligocene mesotheriids (Mammalia, Notoungulata) from Salla and Lacayani (Bolivia): implications for basal mesotheriid phylogeny and distribution", Zoological Journal of the Linnean Society, 152: 153–200, doi:10.1111/j.1096-3642.2007.00388.x, retrieved 2018-09-11

Fray Bentos Formation

• Bond, Mariano; López, Guillermo; Reguero, Marcelo A.; Scillato Yané, Gustavo J.; Vucetich, María G. (1998), "Los mamíferos de la Formación Fray Bentos (Edad mamífero Deseadense, Oligoceno Superior) de las provincias de Corrientes y Entre Ríos, Argentina", Asociación Paleontológica Argentina. Publicación Especial, 5: 41–50, retrieved 2018-09-12
• Tófalo, Ofelia Rita; Morrás, Héctor J.M. (2009), "Evidencias paleoclimáticas en duricostras, paleosuelos y sedimentitas silicoclásticas, del Cenozoico de Uruguay" (PDF), Revista de la Asociación Geológica Argentina: 674–686, retrieved 2018-09-12

Moquegua Formation

• Alván, Aldo; Ramírez, Katerin; von Eynatten, Hilmar; Dunkl, István; Jacay, Javier; Bertone, Gabriela (2017), "Evolución Geológica de las Cuencas de Antearco del Sur de Perú (Moquegua y Camaná-Mollendo): Proveniencia Sedimentaria y Análisis de Facies en Rocas Cenozoicas", Boletín de la Sociedad Geológica del Perú, 112: 53–77, retrieved 2018-09-10
• Shockey, Bruce J.; Billet, Guillaume; Salas Gismondi, Rodolfo (2016), "A new species of Trachytherus (Notoungulata: Mesotheriidae) from the late Oligocene (Deseadan) of Southern Peru and the middle latitude diversification of early diverging mesotheriids", Zootaxa, 4111 (5): 565–583, doi:10.11646/zootaxa.4111.5.3, PMID 27395103, retrieved 2019-02-12

Chambira Formation

• Boivin, Myriam; Marivaux, Laurent; Antoine, Pierre-Olivier (2019), "New insight from the Paleogene record of Amazonia into the early diversification of Caviomorpha (Hystricognathi, Rodentia): phylogenetic, macroevolutionary, and paleobiogeographic implications", Geodiversitas, 41 (4): 143–245, doi:10.5252/geodiversitas2019v41a4, retrieved 2019-02-28
• Boivin, Myriam; Marivaux, Laurent; Candela, Adriana M.; Orliac, Maëva J.; Pujos, François; Salas Gismondi, Rodolfo; Tejada Lara, Julia V.; Antoine, Pierre-Olivier (2016), "Late Oligocene caviomorph rodents from Contamana, Peruvian Amazonia", Papers in Palaeontology, 3 (1): 69–109, doi:10.1002/spp2.1068, hdl:11336/63600, retrieved 2019-02-12
• Marivaux, Laurent; Adnet, Sylvain; Altamirano Sierra, Ali J.; Boivin, Myriam; Pujos, François; Ramdarshan, Anusha; Salas Gismondi, Rodolfo; Tejada, Julia; Antoine, Pierre-Olivier (2016), "Neotropics provide insights into the emergence of New World monkeys: New dental evidence from the late Oligocene of Peruvian Amazonia", Journal of Human Evolution, 97: 159–175, Bibcode:2016JHumE..97..159M, doi:10.1016/j.jhevol.2016.05.011, hdl:11336/182832, PMID 27457552, retrieved 2019-02-04

Barzalosa Formation

• Acosta, Jorge E.; Guatame, Rafael; Caicedo A., Juan Carlos; Cárdenas, Jorge Ignacio (2002), Mapa Geológico de Colombia - Plancha 245 - Girardot - 1:100,000 - Memoria Explicativa, INGEOMINAS, pp. 1–92
• Acosta, Jorge E.; Ulloa, Carlos E. (2001), Mapa Geológico de Colombia - Plancha 246 - Fusagasugá - 1:100,000 - Memoria Explicativa, INGEOMINAS, pp. 1–77

Tremembé Formation

• Do Couto Ribeiro, Graziella (2010), Avaliação morfológica, taxonômica e cronológica dos mamíferos fósseis da Formação Tremembé (Bacia de Taubaté), Estado de São Paulo, Brasil, Universidade de São Paulo, pp. 1–112
• Malabarba, Maria Claudia; Lundberg, John G. (2007), "A fossil loricariid catfish (Siluriformes: Loricarioidea) from the Taubaté Basin, eastern Brazil", Neotropical Ichthyology, 5 (3): 263–270, doi:10.1590/S1679-62252007000300005, retrieved 2018-10-08
• Olson, Storrs L.; Alvarenga, Herculano M.F. (2002), "A new genus of small teratorn from the Middle Tertiary of the Taubate Basin, Brazil (Aves: Teratornithidae)" (PDF), Proceedings of the Biological Society of Washington, 115: 701–705, retrieved 2017-09-04

Las Cascadas Formation

• Rincón, A. F.; Bloch, J. I.; Suárez, C.; MacFadden, B. J.; Jaramillo, C. A. (2012), "New floridatragulines (Mammalia, Camelidae) from the early Miocene Las Cascadas Formation, Panama", Journal of Vertebrate Paleontology, 32 (2): 456–475, Bibcode:2012JVPal..32..456R, doi:10.1080/02724634.2012.635736

Further reading

• Encinas, Alfonso; Folguera, Andrés; Bechis, Florencia; Finger, Kenneth L.; Zambrano, Patricio; Pérez, Felipe; Benarbé, Pablo; Tapia, Francisca; Riffo, Ricardo; Buatois, Luis; Orts, Darío; Nielsen, Sven N.; Valencia, Victor V.; Cituño, José; Oliveros, Verónica; De Girolamo Del Mauro, Lizet; Ramos, Víctor A. (2018). "The Late Oligocene–Early Miocene Marine Transgression of Patagonia". In Folguera, A.; Contreras Reyes, E.; Heredia, N.; et al. (eds.). The Evolution of the Chilean-Argentinean Andes. Springer. pp. 443–474. ISBN 978-3-319-67774-3.
• Woodburne, M.O (2010), "The Great American Biotic Interchange: Dispersals, Tectonics, Climate, Sea Level and Holding Pens", Journal of Mammalian Evolution, 17 (4): 245–264, doi:10.1007/s10914-010-9144-8, PMC 2987556, PMID 21125025
• Webb, S. David (2006), "The Great American Biotic Interchange: Patterns and Processes", Annals of the Missouri Botanical Garden, 93 (2): 245–257, doi:10.3417/0026-6493(2006)93[245:TGABIP]2.0.CO;2