^Vladimir A. Lukhtanov. 2017. A New Species of Melitaea from Israel, with Notes on Taxonomy, Cytogenetics, Phylogeography and Interspecific Hybridization in the Melitaea persea complex (Lepidoptera, Nymphalidae). Comparative Cytogenetics. 11(2); 325-357. DOI: 10.3897/CompCytogen.v11i2.12370
Leneveu, J., Chichvarkhin, A. & Wahlberg, N. 2009 Varying rates of diversification in the genus Melitaea (Lepidoptera: Nymphalidae) during the past 20 million years. Biol. J. Linn. Soc. 97:346-361.
Wahlberg, N., A. V. Z. Brower, and S. Nylin. 2005. Phylogenetic relationships and historical biogeography of tribes and genera in the subfamily Nymphalinae (Lepidoptera: Nymphalidae). Biological Journal of the Linnean Society 86:227-251.
Wahlberg, N., and M. Zimmermann. 2000. Pattern of phylogenetic relationships among members of the tribe Melitaeini (Lepidoptera: Nymphalidae) inferred from mtDNA sequences. Cladistics 16:347-363.