^Wahlberg, N. 2006. That awkward age for butterflies: insights from the age of the butterfly subfamily Nymphalinae. Systematic Biology 55:703-714. (英文)
Blum MJ, Bermingham E, and Dasmahapatra K. 2003. A molecular phylogeny of the neotropical butterfly genus Anartia (Lepidoptera: Nymphalidae). Molecular Phylogenetics and Evolution 26: 46-55.
Dasmahaptra KK, Blum MJ, Aiello A, Hackwell S, Davies N, Bermingham EP, and Mallet J. 2002. Inferences from a rapidly moving hybrid zone. Evolution 56: 741-753.
Lamas G ed. 2004. Atlas of Neotropical Lepidoptera. Checklist: Part 4A Hesperioidea - Papiionoidea. Gainesville: Scientific Publishers/Association of Tropical Lepidoptera.