Genus of ground crab spiders
Female Xysticus sp. with egg sac
Xysticus is a genus of ground crab spiders described by C. L. Koch in 1835, belonging to the order Araneae , family Thomisidae . The genus name is derived from the Ancient Greek root xyst , meaning "scraped, scraper".
Description
Xysticus and Coriarachne are dark brown or reddish-brown crab spiders often encountered on weeds or trees. While similar to the 'flower spiders', they tend to have shorter, sturdier legs. Many, but not all, species have abdomens more patterned than most Thomisus species, rather like some of the Synema species. Some however, that are more terrestrial, resemble earth in colour and texture.
A ground crab spider with earthy coloration
Most species of the genus Xysticus are small to medium sized spiders. They show a sexual dimorphism in size. Females of typical species reach a maximum of 10 millimetres (0.39 in) of body length, while their males are about 3–5 millimetres (0.12–0.20 in) long, about half the size of the females.
The basic colour of these species is usually brown, beige or gray. The prosoma is sometimes slightly smaller than the opisthosoma . In many species the front body shows in the middle a broad, pale longitudinal band. The opisthosoma is clearly flattened in most species and shows a median broad and dark leaf marking.
The species are often very similar to each other, and in most cases can be distinguished only by a microscopic examination of the reproductive organs.
Behaviour
The spiders of the genus Xysticus do not build webs; like most Thomisidae they are ambush hunters and prefer to hunt near the ground (hence the common name "ground crab spiders"). They move slowly, and commonly hunt by stationing themselves in a high-traffic area and grabbing whatever arthropod passes close enough. Also like most other Thomisidae, they seize prey with their enlarged anterior two pairs of legs and kill it by a venomous bite.
Species
As of December 2022[update] it contains 293 species, found worldwide:[ 1]
Xysticus abditus Logunov, 2006 — Bulgaria, Turkey
Xysticus acerbus Thorell, 1872 — Europe to Central Asia
Xysticus acerbus obscurior Kulczyński, 1895 — Ukraine
Xysticus acquiescens Emerton, 1919 — Holarctic
Xysticus advectus O. P.-Cambridge, 1890 — Guatemala, Costa Rica
Xysticus aethiopicus L. Koch, 1875 — Ethiopia
Xysticus albertensis Dondale, 2008 — Canada
Xysticus albolimbatus Hu, 2001 — China
Xysticus alboniger Turnbull, Dondale & Redner, 1965 — USA, Canada
Xysticus aletaiensis Hu & Wu, 1989 — China
Xysticus alpicola Kulczyński, 1882 — Czech Republic, Slovakia, Poland, Ukraine
Xysticus alpinistus Ono, 1978 — Nepal, China
Xysticus alpinus Kulczyński, 1887 — Alps (Switzerland, Italy, Germany, Austria)
Xysticus alsus Song & Wang, 1994 — China
Xysticus altaicus Simon, 1895 — Kazakhstan
Xysticus altitudinis Levy, 1976 — Israel
Xysticus ampullatus Turnbull, Dondale & Redner, 1965 — USA, Canada
Xysticus apachecus Gertsch, 1933 — USA
Xysticus apalacheus Gertsch, 1953 — USA
Xysticus apertus Banks, 1898 — Mexico
Xysticus apricus L. Koch, 1876 — Italy
Xysticus aprilinus Bryant, 1930 — USA
Xysticus arenarius Thorell, 1875 — Ukraine
Xysticus arenicola Simon, 1875 — France
Xysticus argenteus Jézéquel, 1966 — Ivory Coast
Xysticus asper (Lucas, 1838) — Canary Islands
Xysticus atevs Ovtsharenko, 1979 — Russia
Xysticus atrimaculatus Bösenberg & Strand, 1906 — China, Korea, Japan
Xysticus auctificus Keyserling, 1880 — USA, Canada
Xysticus audax (Schrank, 1803) — Palearctic
Xysticus audax massanicus Simon, 1932 — France
Xysticus audaxoides Zhang, Zhang & Song, 2004 — China
Xysticus austrosibiricus Logunov & Marusik, 1998 — Russia, Mongolia, China
Xysticus autumnalis L. Koch, 1875 — New South Wales
Xysticus aztecus Gertsch, 1953 — Mexico
Xysticus bacurianensis Mcheidze, 1971 — Turkey, Russia, Georgia, Azerbaijan
Xysticus bakanas Marusik & Logunov, 1990 — Kazakhstan
Xysticus banksi Bryant, 1933 — USA
Xysticus barbatus Caporiacco, 1936 — Libya
Xysticus benefactor Keyserling, 1880 — USA, Canada
Xysticus bengalensis Tikader & Biswas, 1974 — India
Xysticus bengdakus Saha & Raychaudhuri, 2007 — India
Xysticus beni Strand, 1913 — Central Africa
Xysticus berlandi Schenkel, 1963 — China
Xysticus bermani Marusik, 1994 — Russia, China
Xysticus bharatae Gajbe & Gajbe, 1999 — India
Xysticus bicolor L. Koch, 1867 — Greece
Xysticus bicuspis Keyserling, 1887 — USA
Xysticus bifasciatus C. L. Koch, 1837 — Palearctic
Xysticus bimaculatus L. Koch, 1867 — Queensland
Xysticus bohdanowiczi Zhang, Zhu & Song, 2004 — China
Xysticus bolivari Gertsch, 1953 — Mexico
Xysticus bradti Gertsch, 1953 — Mexico
Xysticus breviceps O. P.-Cambridge, 1885 — India
Xysticus brevidentatus Wunderlich, 1995 — Italy, Croatia, Bosnia and Herzegoniva, Montenegro, Albania
Xysticus britcheri Gertsch, 1934 — Russia, Alaska, Canada, USA
Xysticus brunneitibiis Caporiacco, 1939 — Ethiopia
Xysticus californicus Keyserling, 1880 — USA, Mexico
Xysticus canadensis Gertsch, 1934 — Russia, USA, Canada
Xysticus caspicus Utochkin, 1968 — Russia, Turkmenistan, Kazakhstan
Xysticus caucasius L. Koch, 1878 — Georgia
Xysticus chaparralis Schick, 1965 — USA
Xysticus charitonowi Mcheidze, 1971 — Georgia
Xysticus chippewa Gertsch, 1953 — Holarctic
Xysticus chui Ono, 1992 — Taiwan
Xysticus clercki (Audouin, 1826) — Egypt, Ethiopia
Xysticus cochise Gertsch, 1953 — USA
Xysticus coloradensis Bryant, 1930 — USA
Xysticus concinnus Kroneberg, 1875 — Central Asia
Xysticus concretus Utochkin, 1968 — Russia, China, Korea, Japan
Xysticus concursus Gertsch, 1934 — USA
Xysticus conflatus Song, Tang & Zhu, 1995 — China
Xysticus connectens Kulczyński, 1901 — China
Xysticus cor Canestrini, 1873 — Southern Europe, Turkey, Azores, Iran
Xysticus corsicus Simon, 1875 — Corsica
Xysticus crispabilis Song & Gao, 1996 — China
Xysticus cristatus (Clerck, 1757) — Palearctic, Introduced to Canada, USA
Xysticus croceus Fox, 1937 — India, Nepal, Bhutan, China, Vietnam, Korea, Taiwan, Japan
Xysticus cunctator Thorell, 1877 — USA, Canada
Xysticus curtus Banks, 1898 — Mexico
Xysticus dali Li & Yang, 2008 — China
Xysticus davidi Schenkel, 1963 — Russia, China
Xysticus denisi Schenkel, 1963 — China
Xysticus desidiosus Simon, 1875 — Europe
Xysticus discursans Keyserling, 1880 — North America
Xysticus diversus (Blackwall, 1870) — Sicily
Xysticus doriai (Dalmas, 1922) — Italy
Xysticus dzhungaricus Tyschchenko, 1965 — Russia, Central Asia to China
Xysticus edax (O. P.-Cambridge, 1872) — Turkey, Israel
Xysticus elegans Keyserling, 1880 — USA, Canada, Alaska
Xysticus elephantus Ono, 1978 — Nepal, China
Xysticus ellipticus Turnbull, Dondale & Redner, 1965 — USA, Canada
Xysticus emertoni Keyserling, 1880 — USA, Canada, Alaska, Slovakia to China
Xysticus ephippiatus Simon, 1880 — Kazakhstan, Russia, Central Asia, Mongolia, China, Korea, Japan
Xysticus erraticus (Blackwall, 1834) — Europe, Russia
Xysticus facetus O. P.-Cambridge, 1896 — Mexico to El Salvador
Xysticus fagei Lessert, 1919 — East Africa
Xysticus federalis Gertsch, 1953 — Mexico
Xysticus ferox (Hentz, 1847) — USA, Canada
Xysticus ferrugineus Menge, 1876 — Palearctic
Xysticus ferruginoides Schenkel, 1963 — Russia, Mongolia
Xysticus fervidus Gertsch, 1953 — USA, Canada
Xysticus flavitarsis Simon, 1877 — Congo
Xysticus flavovittatus Keyserling, 1880 — USA
Xysticus floridanus Banks, 1896 — USA
Xysticus fraternus Banks, 1895 — USA, Canada
Xysticus funestus Keyserling, 1880 — North America
Xysticus furtivus Gertsch, 1936 — USA
Xysticus gallicus Simon, 1875 — Palearctic
Xysticus gallicus batumiensis Mcheidze & Utochkin, 1971 — Georgia
Xysticus gattefossei Denis, 1956 — Morocco
Xysticus geometres L. Koch, 1874 — Queensland
Xysticus gertschi Schick, 1965 — North America
Xysticus ghigii Caporiacco, 1938 — Mexico
Xysticus gortanii Caporiacco, 1922 — Italy
Xysticus gosiutus Gertsch, 1933 — USA, Canada
Xysticus gracilis Keyserling, 1880 — Colombia
Xysticus grallator Simon, 1932 — Portugal, Spain, France (Corsica), Italy (Sardinia)
Xysticus guizhou Song & Zhu, 1997 — China
Xysticus gulosus Keyserling, 1880 — North America
Xysticus havilandi Lawrence, 1942 — South Africa
Xysticus hedini Schenkel, 1936 — Russia, Mongolia, China, Korea, Japan
Xysticus helophilus Simon, 1890 — Yemen
Xysticus hepaticus Simon, 1903 — Madagascar
Xysticus himalayaensis Tikader & Biswas, 1974 — India
Xysticus hindusthanicus Basu, 1965 — India
Xysticus hui Platnick, 1993 — China
Xysticus humilis Redner & Dondale, 1965 — USA
Xysticus ibex Simon, 1875 — France, Spain
Xysticus ibex dalmasi Simon, 1932 — France
Xysticus ictericus L. Koch, 1874 — Fiji
Xysticus idolothytus Logunov, 1995 — Kazakhstan, Mongolia
Xysticus illaudatus Logunov, 1995 — Russia
Xysticus imitarius Gertsch, 1953 — USA
Xysticus indiligens (Walckenaer, 1837) — USA
Xysticus insulicola Bösenberg & Strand, 1906 — China, Korea, Japan
Xysticus iviei Schick, 1965 — USA
Xysticus iviei sierrensis Schick, 1965 — USA
Xysticus jabalpurensis Gajbe & Gajbe, 1999 — India
Xysticus jaharai Basu, 1979 — India
Xysticus japenus Roewer, 1938 — Indonesia
Xysticus jiangi Peng, Yin & Kim, 2000 — China
Xysticus jinlin Song & Zhu, 1995 — China
Xysticus joyantius Tikader, 1966 — India
Xysticus jugalis L. Koch, 1875 — Ethiopia
Xysticus jugalis larvatus Caporiacco, 1949 — Kenya
Xysticus kalandadzei Mcheidze & Utochkin, 1971 — Georgia
Xysticus kali Tikader & Biswas, 1974 — India
Xysticus kamakhyai Tikader, 1962 — India
Xysticus kansuensis Tang, Song & Zhu, 1995 — China, Japan
Xysticus kashidi Tikader, 1963 — India
Xysticus kaznakovi Utochkin, 1968 — Greece to Central Asia
Xysticus kempeleni Thorell, 1872 — Europe to Central Asia
Xysticus kempeleni nigriceps Simon, 1932 — France
Xysticus khasiensis Tikader, 1980 — India
Xysticus kochi Thorell, 1872 — Europe, Mediterranean to Central Asia
Xysticus kochi abchasicus Mcheidze & Utochkin, 1971 — Georgia
Xysticus kulczynskii Wierzbicki, 1902 — Azerbaijan, Iran
Xysticus kurilensis Strand, 1907 — Russia, China, Korea, Japan
Xysticus kuzgi Marusik & Logunov, 1990 — Central Asia
Xysticus laetus Thorell, 1875 — Italy to Central Asia
Xysticus lanio C. L. Koch, 1835 — Palearctic
Xysticus lanio alpinus Kulczyński, 1887 — Austria
Xysticus lapidarius Utochkin, 1968 — Central Asia
Xysticus lassanus Chamberlin, 1925 — USA, Mexico
Xysticus latitabundus Logunov, 1995 — Russia
Xysticus lendli Kulczyński, 1897 — Hungary
Xysticus lepnevae Utochkin, 1968 — Russia, Korea, Sakhalin
Xysticus lineatus (Westring, 1851) — Palearctic
Xysticus locuples Keyserling, 1880 — North America
Xysticus logunovi Seyfulina & Mikhailov, 2004 — Russia
Xysticus logunovorum Blick & Ono, 2021 — Iran
Xysticus lucifugus Lawrence, 1937 — South Africa
Xysticus luctans (C. L. Koch, 1845) — USA, Canada
Xysticus luctator L. Koch, 1870 — Palearctic
Xysticus luctuosus (Blackwall, 1836) — Holarctic
Xysticus lutzi Gertsch, 1935 — USA, Mexico
Xysticus macedonicus Silhavy, 1944 — Germany, Switzerland, Austria, Italy, Albania, North Macedonia, Greece, Turkey
Xysticus maculatipes Roewer, 1962 — Afghanistan
Xysticus maculiger Roewer, 1951 — Yarkand
Xysticus manas Song & Zhu, 1995 — China
Xysticus marmoratus Thorell, 1875 — Hungary, Slovakia, Bulgaria, Russia, Ukraine
Xysticus martensi Ono, 1978 — Nepal
Xysticus marusiki Ono & Martens, 2005 — Turkey, Iran
Xysticus metinaktasi Demir, Seyyar & Türker, 2017 — Turkey
Xysticus montanensis Keyserling, 1887 — USA, Canada, Alaska
Xysticus mugur Marusik, 1990 — Russia
Xysticus mulleri Lawrence, 1952 — South Africa
Xysticus multiaculeatus Caporiacco, 1940 — Ethiopia
Xysticus mundulus O. P.-Cambridge, 1885 — Yarkand
Xysticus namaquensis Simon, 1910 — South Africa
Xysticus natalensis Lawrence, 1938 — South Africa
Xysticus nataliae Utochkin, 1968 — Russia
Xysticus nigriceps Berland, 1922 — East Africa
Xysticus nigropunctatus L. Koch, 1867 — Queensland
Xysticus nigrotrivittatus (Simon, 1870) — Portugal, Spain
Xysticus nitidus Hu, 2001 — China
Xysticus nubilus Simon, 1875 — Macaronesia, Mediterranean, Georgia
Xysticus nyingchiensis Song & Zhu, 1995 — China
Xysticus obscurus Collett, 1877 — Holarctic
Xysticus ocala Gertsch, 1953 — USA
Xysticus orizaba Banks, 1898 — Mexico
Xysticus ovatus Simon, 1876 — France
Xysticus paiutus Gertsch, 1933 — USA, Mexico
Xysticus palawanicus Barrion & Litsinger, 1995 — Philippines
Xysticus palpimirabilis Marusik & Chevrizov, 1990 — Kyrgyzstan
Xysticus parallelus Simon, 1873 — Corsica, Sardinia
Xysticus parapunctatus Song & Zhu, 1995 — China
Xysticus pearcei Schick, 1965 — USA, Mexico
Xysticus peccans O. P.-Cambridge, 1876 — Egypt
Xysticus pellax O. P.-Cambridge, 1894 — North America
Xysticus peltiformus Zhang, Zhu & Song, 2004 — China
Xysticus peninsulanus Gertsch, 1934 — USA
Xysticus periscelis Simon, 1908 — Western Australia
Xysticus pieperi Ono & Martens, 2005 — Iran
Xysticus pigrides Mello-Leitao, 1929 — Cape Verde Islands
Xysticus pingshan Zhang, Zhu & Song, 2004 — China
Xysticus posti Sauer, 1968 — USA
Xysticus pretiosus Gertsch, 1934 — USA, Canada
Xysticus promiscuus O. P.-Cambridge, 1876 — Egypt, Israel
Xysticus pseudobliteus (Simon, 1880) — Russia, Kazakhstan, Mongolia, China, Korea
Xysticus pseudocristatus Azarkina & Logunov, 2001 — Pakistan, Central Asia to China
Xysticus pseudolanio Wunderlich, 1995 — Bulgaria, Turkey, Georgia
Xysticus pseudoluctuosus Marusik & Logunov, 1995 — Turkey, Tajikistan
Xysticus pulcherrimus Keyserling, 1880 — Colombia
Xysticus punctatus Keyserling, 1880 — USA, Canada
Xysticus pygmaeus Tyschchenko, 1965 — Kazakhstan
Xysticus pynurus Tikader, 1968 — India
Xysticus quadrispinus Caporiacco, 1933 — Libya
Xysticus quadrispinus concolor Caporiacco, 1933 — Libya
Xysticus quagga Jocque, 1977 — Morocco
Xysticus rainbowi Strand, 1901 — New Guinea
Xysticus robinsoni Gertsch, 1953 — USA, Mexico
Xysticus rockefelleri Gertsch, 1953 — Mexico
Xysticus roonwali Tikader, 1964 — India, Nepal
Xysticus rostratus Ono, 1988 — Russia, Japan
Xysticus ryukyuensis Ono, 2002 — Ryukyu Islands
Xysticus saganus Bösenberg & Strand, 1906 — Russia, China, Korea, Japan
Xysticus sagittifer Lawrence, 1927 — Namibia
Xysticus sansan Levy, 2007 — Israel
Xysticus schoutedeni Lessert, 1943 — Congo
Xysticus semicarinatus Simon, 1932 — France, Spain, Portugal
Xysticus sharlaa Marusik & Logunov, 2002 — Russia, China
Xysticus shillongensis Tikader, 1962 — India
Xysticus shyamrupus Tikader, 1966 — India
Xysticus siciliensis Wunderlich, 1995 — Sicily
Xysticus sicus Fox, 1937 — Russia, China, Korea
Xysticus sikkimus Tikader, 1970 — India, China
Xysticus silvestrii Simon, 1905 — Argentina
Xysticus simonstownensis Strand, 1909 — South Africa
Xysticus sjostedti Schenkel, 1936 — Russia, Kazakhstan, Mongolia, China
Xysticus slovacus Svaton, Pekar & Prídavka, 2000 — Slovakia, Russia
Xysticus soderbomi Schenkel, 1936 — Mongolia, China
Xysticus soldatovi Utochkin, 1968 — Russia, China
Xysticus spasskyi Utochkin, 1968 — Ukraine, Russia, Georgia, Azerbaijan
Xysticus sphericus (Walckenaer, 1837) — USA
Xysticus spiethi Gertsch, 1953 — Mexico
Xysticus subjugalis Strand, 1906 — Ethiopia
Xysticus subjugalis nigerrimus Caporiacco, 1941 — Ethiopia
Xysticus tampa Gertsch, 1953 — USA
Xysticus tarcos L. Koch, 1875 — Ethiopia
Xysticus taukumkurt Marusik & Logunov, 1990 — Kazakhstan
Xysticus tenuiapicalis Demir, 2012 — Turkey
Xysticus texanus Banks, 1904 — USA, Mexico
Xysticus thessalicoides Wunderlich, 1995 — Greece, Crete, Turkey
Xysticus thessalicus Simon, 1916 — Albania, Bulgaria, Croatia, Greece, Turkey, Israel[ 2]
Xysticus tikaderi Bhandari & Gajbe, 2001 — India
Xysticus toltecus Gertsch, 1953 — Mexico
Xysticus torsivoides Song & Zhu, 1995 — China
Xysticus tortuosus Simon, 1932 — Portugal to Austria, Morocco, Algeria
Xysticus transversomaculatus Bösenberg & Strand, 1906 — Korea, Japan
Xysticus triguttatus Keyserling, 1880 — USA, Canada
Xysticus trizonatus Ono, 1988 — Japan
Xysticus tsanghoensis Hu, 2001 — China
Xysticus tugelanus Lawrence, 1942 — South Africa
Xysticus turkmenicus Marusik & Logunov, 1995 — Central Asia
Xysticus ukrainicus Utochkin, 1968 — Russia, Georgia
Xysticus ulmi (Hahn, 1831) — Palearctic
Xysticus urbensis Lawrence, 1952 — South Africa
Xysticus urgumchak Marusik & Logunov, 1990 — Central Asia
Xysticus variabilis Keyserling, 1880 — USA
Xysticus verecundus Gertsch, 1934 — Mexico
Xysticus verneaui Simon, 1883 — Canary Islands, Madeira
Xysticus viduus Kulczyński, 1898 — Palearctic
Xysticus viveki Gajbe, 2005 — India
Xysticus wagneri Gertsch, 1953 — Mexico
Xysticus walesianus Karsch, 1878 — New South Wales
Xysticus winnipegensis Turnbull, Dondale & Redner, 1965 — Canada
Xysticus wunderlichi Logunov, Marusik & Trilikauskas, 2001 — Russia, China
Xysticus xerodermus Strand, 1913 — Turkey, Israel
Xysticus xiningensis Hu, 2001 — China
Xysticus yebongensis Lee & Kim, 2018 — Korea
Xysticus yogeshi Gajbe, 2005 — India
Gallery
Xysticus audax
Xysticus kochi , female
Xysticus cristatus
Xysticus bilimbatus
Xysticus lanio
Xysticus luctator
Xysticus ninniii
Xysticus ulmi
References
Norman I. Platnick - Thomisidae
J. H. Redner and C. D. Dondale (1965). A New Species of The Spider Genus Xysticus (Araneae: Thomisidae) From Arizona - Psyche 72(4): 291–294.
H. Demir, O. Seyyar, and M. Aktaş (2008). A Poorly Known Species of the Spider Genus Xysticus C. L. Koch (Araneae, Thomisidae) in Turkey. Archives of Biological Sciences 60(4): 17P-18P.
External links