^Morcillo, F.; Ornelas-García, C.P.; Alcaraz, L.; Matamoros, W.A. & Doadrio, I. (2015). "Phylogenetic relationships and evolutionary history of the Mesoamerican endemic freshwater fish family Profundulidae (Cyprinodontiformes: Actinopterygii)". Molecular Phylogenetics and Evolution. 94 (Part. A): 242–251. doi:10.1016/j.ympev.2015.09.002. PMID26364972.
^Dominguez-Cisneros, Sara E, Ernesto Velázquez-Velázquez, Caleb D. McMahan, Wilfredo A. Matamoros; A new species of killifish of the genus Profundulus (Atherinomorpha: Profundulidae) from the upper reaches of the Papaloapan River in the Mexican state of Oaxaca. 2021. Ichthyology & Herpetology. 109(4):949–957
^Luis Fernando Del Moral-Flores, Eduardo López-Segovia & Tao Hernández-Arellano; Profundulus chimalapensis, una nueva especie de pez ciprinodóntido (Cyprinodontiformes: Profundulidae) del Río Coatzacoalcos, México | https://revistas.ucr.ac.cr/index.php/rbt/article/view/40340/44229