P-22

This article is about the mountain lion. For other uses, see P22.

P-22

P-22 in 2019
Other name(s)	P-022[1] Puma 22 Hollywood Cat
Species	Cougar (Puma concolor)
Sex	Male
Born	c. 2009 or 2010[1] Santa Monica Mountains, California, U.S.
Died	December 17, 2022(2022-12-17) (aged 12–13) San Diego, California, U.S.[2]
Cause of death	Euthanasia
Resting place	Griffith Park, Los Angeles, California
Residence	Griffith Park, Los Angeles, California
Parent(s)	P-001 (father) unknown female mountain lion (mother)
Weight	123 lb (56 kg) (2012)[1]

P-22 (c. 2009/2010 – December 17, 2022) was a wild mountain lion who resided in Griffith Park in Los Angeles, California, on the eastern side of the Santa Monica Mountains.[1][3] P-22 was first identified in 2012[4] and was the subject of significant media attention, including numerous books, television programs and other works of art.[5][6] P-22 was often recorded prowling in the Hollywood Hills neighborhood of Los Angeles.[6][7][8] He was monitored by a radio collar.[9][10] P-22 remained in Griffith Park for ten years until he was captured on December 12, 2022. He was then euthanized on December 17, 2022, after examinations revealed he was suffering from traumatic injuries consistent with being hit by a car, in combination with several longer-term health issues.[11][12]

Early life

P-22 was born circa 2010 in the western part of the Santa Monica Mountains.[1] Genetic testing showed that his father was P-1.[13] P-1 was known as "king of the mountains," having a very large territory.[14] His mother was an unknown puma, one who had not been recorded in the study of local pumas.[1]

Sometime before 2012, P-22 headed east within the Santa Monica Mountains to Griffith Park, where he settled after crossing two major Los Angeles freeways (Interstate 405 and Route 101).[3][15] His success in evading traffic on these major routes was highlighted, since multiple mountain lions have died after being struck by vehicles on Los Angeles freeways.[6][16] The exact route for P-22's journey is unknown.[17]

In Los Angeles

P-22's home range in comparison to other pumas'.

The U.S. National Park Service noted that P-22's nine-square-mile Griffith Park habitat (23 km^2) is too small for an adult cat by a factor of 31 and that it was unlikely he would ever find a mate there.[1] It’s the smallest ever recorded range for an adult male mountain lion.[18][note 1]

P-22 primarily resided in Griffith Park but was spotted by Angelenos throughout the Los Angeles neighborhoods of the Hollywood Hills,[20][21][22][23] Los Feliz,[24][25] and Silver Lake.[26][27]

2012 discovery

The Griffith Park Connectivity Study, funded by Friends of Griffith Park, was launched in mid-2011. FoGP entered a study agreement and purchased thirteen wildlife cameras. The study objective was to evaluate the movement of large and medium-sized mammals to and from Griffith Park and the surrounding open space. It was the first project in the Griffith Park region targeting potential corridors. The study was expanded further in 2013, to include tunnels to the LA River and other passages.

Daniel S. Cooper led the study. Erin Boydston of USGS Western Ecological Research Center also supported the project. Miguel Ordeñana joined as a field biologist. The three ecologists, Cooper, Boydston, and Ordeñana, soon began to document deer, bobcats and coyotes crossing via one of the overpass bridges of the Hollywood Freeway in Cahuenga Pass. Ordeñana, poring over hundreds of motion-triggered photos, saw the first camera image taken on February 12, 2012 of the hind quarters of a male mountain lion on a rugged ridgeline just above Ford Theatre. The team released a Study Update in March 2012.[28][29][30][31]

Jeff Sikich led the mission to catch him. He was first caught in March of 2012.[29] He was fitted with an electronic neck collar that recorded his location over time.[17] He then weighed 90 pounds (41 kg).[1] He was designated P-22.[29] P is short for "puma" and 22 refers to him being the 22nd puma in the ongoing puma study.[1][3][note 2]

2013 National Geographic photos

National Geographic photographer Steve Winter worked with Jeff Sikich, a wildlife biologist with the National Park Service,[33][35] to photograph P-22. He spent 15 months putting up camera traps in Griffith Park, and getting his cameras stolen, before capturing the now-famous photo of P-22 under the Hollywood Sign.[9][35] This image appeared in the December 2013 issue of National Geographic.[35]

2014 health issues

P-22 in March 2014 before his treatment for mange (left) and in December 2015, after treatment (right)

In 2014, the National Park Service reported that P-22 had contracted mange stemming from exposure to anti-blood-clotting rat poison by way of biomagnification.[36][37] The Park Service captured P-22, administered topical medications and injections of vitamin K in efforts to treat him, then released him back into Griffith Park. The mange eventually subsided and P-22's health improved. The National Park Service again captured P-22 in December 2015 and found he had fully recovered, gaining 15 pounds.[38][39]

2016 koala killing

In 2016, the Los Angeles Zoo reported the disappearance of an elderly koala named Killarney, whose carcass was found outside of the koala enclosure.[40][41][42] Surveillance footage from the zoo showed P-22 (then seven years old) nearby on zoo grounds, although neither the GPS-tracking data nor camera footage recorded an actual interaction between the two animals.[40][41][43] Los Angeles City Council member Mitch O'Farrell called for investigating the relocation of P-22 after the incident.[44] The National Park Service called the koala killing "normal predatory behavior", and the Zoo declined to ask for a depredation permit for P-22, instead opting for more secure enclosure methods for some of its animals at night.[42]

Capture and death

The National Park Service and the California Department of Fish and Wildlife announced in December 2022 that they would capture P-22 to evaluate his health,[45] following a pair of attacks on chihuahuas in the city.[46] Changes in his behavior had also been noticed including appearing to be agitated and venturing farther from his usual range.[47] On December 12, he was located in a Los Feliz homeowner's backyard and tranquilized by the California Department of Fish and Wildlife.[48] He was first triaged at the Los Angeles Zoo, and then taken to the San Diego Zoo Wildlife Alliance. Officials initially said he was in stable condition[48][49] and were considering all options, including releasing P-22 back into the wild or keeping him in a wildlife sanctuary, but that euthanasia was off the table unless P-22 was determined to be suffering from a "really serious" health condition.[50]

The next day, officials announced that P-22 would likely not be released back into Griffith Park, and that his health had deteriorated. According to the Department of Fish and Wildlife and the Park Service, he was found to be significantly underweight, with thinning fur, possible mange, and damage to his right eye, possibly from a vehicle—a vehicle collision with a mountain lion had been reported the night of December 11, and P-22's radio collar data supported the idea that he had been the mountain lion involved.[11][51] Officials said P-22 would undergo further medical evaluations before a decision was made.[51]

He was examined by a team of doctors at the San Diego Zoo Safari Park which revealed serious health issues. His injuries included skull fractures, injuries to his right eye, skin injuries, and herniation of abdominal organs into his chest.[52] In addition to those health issues, P-22 was found to be suffering from multiple longer-term medical problems, including stage 2 kidney failure, heart disease, a parasitic skin infection of Demodex gatoi, and weight loss (he weighed 90 lb (41 kg) instead of his typical 125 lb (57 kg)).[12][2][53] On December 17, at 9:00 a.m., P-22 was euthanized.[11][12] A necropsy found he had a systemic ringworm infection which was a first to find this concurrent with a demodectic mange infection in a California mountain lion. The examination confirmed that the multiple severe injuries and chronic conditions impaired his ability to function in the wild and would have lowered his quality of life if placed in human care.[54]

Burial

More than a year prior to P-22's actual death, Miguel Ordeñana applied for the Natural History Museum — in the event of P-22's death — to receive his remains for research purposes, and thought that P-22 would be put on display at the museum.[55] When P-22 died, local Native American tribes, who call mountain lions teachers, requested P-22 to be buried near Griffith Park with a ceremony that honors his spirit.[56] The museum announced that they did not plan to taxidermy his body or put his remains on display.[55]

After a necropsy, P-22 was transported from San Diego to the Natural History Museum of Los Angeles County.[12] A blessing ceremony was held by museum officials and descendants from Gabrieleño/Tongva, Tataviam, and Chumash, Gabrielino-Shoshone, Akimel O'otham, and Luiseño Tribes to "welcome P-22 back to his homeland".[57] His remains were stored in a freezer at the museum until he was buried on March 4 in the Santa Monica Mountains at an undisclosed location.[58] The funeral was done in collaboration with local Indigenous partners, the California Department of Fish and Wildlife, the Natural History Museum and the National Park Service.[59][60][61]

Legacy and tributes

Park rangers and volunteers during P-22 Day in 2017

Numerous members of the community and public figures released statements or condolences shortly following news of his death, including Governor of California Gavin Newsom, U.S. representative Adam Schiff, California State Representative Laura Friedman, and Los Angeles City Councilmember Nithya Raman, among many others.[62][63][64][65][66] In his statement, Newsom said that "P-22's survival on an island of wilderness in the heart of Los Angeles captivated people around the world and revitalized efforts to protect our diverse native species and ecosystems."[62] The Greek Theatre in Griffith Park was the site of a celebration of the life of P-22, presented by National Wildlife Federation on February 4, 2023.[67]

In 2016, Beth Pratt, a California regional executive director for the National Wildlife Federation helped establish the #SaveLACougars campaign and held the first celebration of P-22 Day.[68] The Natural History Museum of Los Angeles County has an exhibit about him.[69]

Since first being spotted in Los Angeles, P-22 became the subject of multiple books, television programs, and works of art. More broadly, the image of P-22 has risen to use as a symbol representing Los Angeles culture and wildlife conservation both in California and more generally.[70][71][68] Articles commemorating P-22 before and after his death noted the challenges the mountain lion faced. The Los Angeles Times' obituary for P-22 called him "an aging bachelor who adjusted to a too-small space in the big city," and described his border-crossing journey to Griffith Park as something that many Los Angeles residents could empathize with.[11]

The Los Angeles Public Library issued a limited-edition library card featuring P-22 in the National Geographic photo with the Hollywood Sign in the background.[72]

In February 2023, Southern California U.S. representatives Adam Schiff, Julia Brownley, and Ted Lieu wrote a letter to the Citizens' Stamp Advisory Committee nominating P-22 to appear on a future stamp design.[73]

Wallis Annenberg Wildlife Crossing

P-22 became the poster puma for the promotion of the Wallis Annenberg Wildlife Crossing. In order to get to Griffith Park, P-22 had to cross two freeways, the 101 and the 405.[74] While he survived his crossings, many pumas do not. The wildlife crossing over the 101 freeway was proposed in 2015. The funds were raised, and the project broke ground in April 2022.[75][76]

Murals

Multiple murals around the city prominently feature P-22.

Muralist	Location	Date painted	Ref
Jonathan Martinez	Esperanza Elementary School in Westlake, Los Angeles	August 2020	[77]
Jonathan Martinez	Watts, Los Angeles	April 2021	[78][79]
Jonathan Martinez	Ladera STARS Academy in Thousand Oaks	December 2022	[80]
Corie Mattie	Silver Lake, Los Angeles	October 2022	[81][82]
Corie Mattie	Fairfax District, Los Angeles	December 2022	[83]
Corie Mattie	Hollywood Boulevard in Hollywood	October 2024	[84]

In popular culture

Books

Year	Title	Author	Publisher	Notes	Ref.
2017	We Heart P-22: A Coloring + Activity Book Celebrating L.A.'s Most Famous Mountain Lion		Narrated Objects		[85][86]
2018	P-22: The Journey	Sherry Mangel-Ferber and Calandra Cherry	Ghost Cat Publications		[87]
2020	The Cat That Changed America	Tony Lee Moral			[5][88]
2020	P-22: The Park	Calandra Cherry and Sherry Mangel-Ferber	Ghost Cat Publications		[5]
2021	Cougar Crossing: How Hollywood's Celebrity Cougar Helped Build a Bridge for City Wildlife	Meeg Pincus	Beach Lane Books	Illustrated by Alexander Vidal	[89][90][91]
2023	Open Throat	Henry Hoke	Farrar, Straus and Giroux	Fictionalized portrayal of P-22

Film and television

Year	Title	Notes	Ref.
2017	The Secret Diary of P-22	Short film	[92][93]
2017	The Cat That Changed America		[71][32][94]
2022	America's Most Infamous Mountain Lion		[29]

P-22's highway journey was parodied in an easter egg in season 4 episode 6 of the television show Bojack Horseman.[95][96] He was also featured in a clue on the game show Jeopardy! in 2022.[97] P-22 was mentioned in the Hulu original series, This Fool. When the main character Julio Lopez is taken for an unwanted hiking trip by his ex-girlfriend Maggie, he worries that they will be devoured by P-22. The mountain lion later appears instead to menace a happy couple who has just gotten engaged.[98][99] A memorial dedication was included at the end of the season 2 premiere episode The Rooster.[100]

Playwright Amy Raasch portrays P-22 in her recurring one-woman stage production The Animal Monologues.[101][102]

Music

• Nick Cave indirectly references P-22 in "Hollywood", the closing track of his 2019 album Ghosteen.[103]

See also

• Fauna of the Santa Monica Mountains
• Cougar–human interactions

Notes

1. P-22 is likely not the first mountain lion to have taken up residence in Griffith Park, although the duration of his stay was remarkably long. A mountain lion's body was found in Griffith Park sometime in 1996 or 1997, after being hit by a vehicle. Another mountain lion was sighted several times in Griffith Park in 2004 and rangers found evidence (including deer remains) to support its presence there.[19]
2. Mountain lions studied by the National Park Service in the Santa Monica Mountains are numbered sequentially by capture date.[32][33] In 2015, radio station KPCC held a poll to choose a "more personal" common-use name for P-22. Proposed names included Felix (like Felix the Cat), Yossarian (based on Catch-22), Tukuurot (Tongva word for mountain lion), Pete Puma (Looney Tunes), Pounce de Leon (Juan Ponce de León), Huell, and Puma Thurman (Uma Thurman). However, the winning name by a large margin was P-22, and so his name stayed.[34]

References

1. "Puma Profiles: P-022". National Park Service. Archived from the original on April 24, 2022. Retrieved October 27, 2022.
2. "Mountain Lion P-22 Compassionately Euthanized Following Complete Health Evaluation Results" (Press release). California Department of Fish and Wildlife. December 17, 2022. Archived from the original on December 26, 2022.
3. Groves, Martha (August 14, 2012). "Mountain lion makes itself at home in Griffith Park". Los Angeles Times. Retrieved October 27, 2022.
4. "Mountain Lion Captured, Collared and Released in Griffith Park Area". National Park Service. April 9, 2012. Retrieved October 27, 2022.
5. Guldimann, Suzanne (December 18, 2020). "Two New Books Celebrate the Life of Mountain Lion P-22". Topanga New Times. Retrieved October 27, 2022.
6. Gammon, Katharine (February 13, 2022). "'The Brad Pitt of mountain lions': how P22 became Los Angeles' wildest celebrity". The Guardian. Retrieved October 27, 2022.
7. Martinez, Christian (April 7, 2022). "Mountain lion P-22 makes another curious visit to the 'luckiest person in L.A.'". Los Angeles Times. Retrieved October 27, 2022.
8. Pridgen, Andrew (October 5, 2022). "'They're out there': Why California mountain lion sightings are on the rise". SFGATE.
9. Keefe, Alexa (November 14, 2013). "A Cougar Ready for His Closeup". National Geographic. Archived from the original on March 1, 2021.
10. Bradley, Ryan (2016). "Mysterious American Cat: The Mountain Lions of Los Angeles". The Virginia Quarterly Review. 92 (3): 24–29. JSTOR 26447081.
11. Nelson, Laura J.; Queally, James (December 17, 2022). "P-22, L.A. celebrity mountain lion, euthanized due to severe injuries". Los Angeles Times. Archived from the original on December 17, 2022. Retrieved December 17, 2022.
12. Kiszla, Cameron (December 17, 2022). "P-22 euthanized days after capture; cougar had been struck by car, wildlife officials say". KTLA. Retrieved December 17, 2022.
13. Nelson, Laura J. (April 21, 2022). "He's terminally single and getting old. What's next for P-22, L.A's favorite wild bachelor?". LA Times. Archived from the original on April 21, 2022.
14. "Life and Death of P-22, L.A.'s Most Famous Mountain Lion". Los Angeles Almanac.
15. Witt, Emily (May 17, 2022). "An Urban Wildlife Bridge is Coming to California". The New Yorker.
16. Goldberg, Noah (August 26, 2022). "Mountain lion P-22 spotted roaming Hollywood Hills streets". Los Angeles Times.
17. Curwen, Thomas (February 8, 2017). "A week in the life of P‑22, the big cat who shares Griffith Park with millions of people". Los Angeles Times. Retrieved October 27, 2022.
18. "Statement on P-22 from Santa Monica Mountains National Recreation Area". National Park Service. December 17, 2022. Retrieved December 17, 2022.
19. Hymon, Steve; Sciaudone, Christiana (April 29, 2004). "A Mountain Lion Far From Home". Los Angeles Times. Retrieved December 18, 2022.
20. Groves, Martha (October 4, 2013). "Scientists track cougar's wild nightlife above Hollywood". Los Angeles Times.
21. Fields, Kayle (March 12, 2014). "Mountain Lion Calls Hollywood Hills Home". ABC News.
22. Solis, Nathan; Yee, Gregory (March 10, 2022). "P-22 is back home in Griffith Park. Here's how he survived urban ills to become L.A.'s most famous cat". Los Angeles Times.
23. Park, Jeong (October 9, 2022). "Los Feliz couple encounter surprise in their driveway: mountain lion believed to be P-22". Los Angeles Times.
24. "Mountain lion P-22 trapped under Los Feliz home". ABC 7 – Eyewitness News. April 13, 2015.
25. Groves, Martha; Jennings, Angel (April 13, 2015). "P-22 vacates home, heads back to Griffith Park, wildlife officials say". Los Angeles Times.
26. Kurland, Zoe (March 10, 2022). "We Saw A Mountain Lion Walking In Silver Lake. The Famous P-22 Takes A Stroll, Then Heads Home To Griffith Park". LAist.
27. Haring, Bruce (March 27, 2022). "Celebrity Mountain Lion P-22 Visits Silver Lake Once Again This Weekend". Deadline.
28. Cooper, Dan; Ordeñana, Miguel; Boydston, Erin (March 2012). Griffith Park Wildlife Connectivity Study Project Update (PDF) (Report). Friends of Griffith Park.
29. "America's Most Infamous Mountain Lion (P22 Puma Documentary)". YouTube. Real Wild. July 9, 2022.
30. Pratt, Beth (2014). "A Mountain Lion in Hollywoodland". When Mountain Lions Are Neighbors (PDF).
31. "P-22's Decade in Griffith Park". Friends of Griffith Park.
32. "P-22's Decade in Griffith Park". Friends of Griffith Park.
33. Nyce, Caroline Mimbs (November 29, 2022). "Tracking the Mountain Lion That Ate a Chihuahua". The Atlantic.
34. Rabe, John (April 24, 2015). "And P-22 the mountain lion's 'new' name, selected by listeners, is..." KPCC.
35. Dell'Amore, Christine (December 14, 2022). "How this photo turned a reclusive mountain lion into a Hollywood icon". National Geographic. Archived from the original on December 14, 2022.
36. "Griffith Park Mountain Lion Exposed to Poison, Suffering from Mange". Santa Monica Mountains National Recreation Area. National Park Service. April 17, 2017. Retrieved October 27, 2022.
37. Hillard, Gloria (June 21, 2014). "LA Mountain Lion A Poster Cat For California's Rat Poison Problem". Weekend Edition Saturday. NPR.
38. Kim, Jed (January 7, 2016). "Mountain lion P-22's health much better in latest checkup". KPCC.
39. Branson-Potts, Hailey (January 7, 2016). "Griffith Park mountain lion P-22 looking healthy again". Los Angeles Times. Archived from the original on December 12, 2022.
40. Martinez, Michael (March 10, 2016). "Mountain lion featured in National Geographic mauls koala in L.A. Zoo". CNN.
41. Serna, Joseph; Branson-Potts, Hailey (March 11, 2016). "Is P-22 mountain lion too dangerous for Griffith Park? Koala death sparks debate". The Baltimore Sun.
42. Kaplan, Sarah (March 17, 2016). "L.A. Zoo to the mountain lion that probably ate its koala: No hard feelings". The Washington Post.
43. Domonoske, Camila (March 11, 2016). "LA's Famous Mountain Lion Suspected In Koala Killing". The Two Way. NPR.
44. Lovett, Ian (March 23, 2016). "Prime Suspect in Koala's Murder: Los Angeles's Mountain Lion". The New York Times. Retrieved December 17, 2022.
45. "California Department of Fish and Wildlife and National Park Service Team Up to Evaluate P-22". wildlife.ca.gov (Press release). California Department of Fish and Wildlife. December 8, 2022. Archived from the original on January 29, 2023. Retrieved December 9, 2022.
46. Yee, Gregory (December 10, 2022). "The search for P-22 is on. Here's why capturing L.A.'s star mountain lion could take weeks". Los Angeles Times.
47. Kranking, Carlyn (December 19, 2022). "Why Los Angeles Fell in Love With the Mountain Lion Known as P-22". Smithsonian Magazine. Retrieved December 28, 2022.
48. Martinez, Christian; Nelson, Laura J.; Solis, Nathan (December 12, 2022). "P-22 captured in backyard of Los Feliz home, resident says". Los Angeles Times. Retrieved December 12, 2022.
49. Albeck-Ripka, Livia (December 12, 2022). "The Lion Sleeps Tonight: P-22, Elusive L.A. Fixture, Is Captured". The New York Times. Retrieved December 13, 2022.
50. Martinez, Christian; Solis, Nathan; Nelson, Laura J.; Yee, Gregory (December 13, 2022). "What will happen to P-22 after his capture? 'No options are off the table' for big cat". Los Angeles Times. Archived from the original on December 13, 2022. Retrieved December 13, 2022.
51. Martinez, Christian; Solis, Nathan (December 13, 2022). "P-22's health seriously deteriorating, with euthanasia or sanctuary possible". Los Angeles Times. Archived from the original on December 14, 2022. Retrieved December 14, 2022.
52. Heyward, Giulia (December 17, 2022). "P-22, Hollywood's famous mountain lion, is euthanized after suffering injuries". NPR News. Retrieved December 22, 2022.
53. Pratt, Beth (December 17, 2022). "A Eulogy for P-22, A Mountain Lion Who Changed the World". nwf.org. National Wildlife Federation.
54. "Final necropsy results released for mountain lion P-22". CDFW News. California Department of Fish and Wildlife. June 14, 2023. Archived from the original on June 14, 2023. Retrieved June 15, 2023.
55. Solis, Nathan (December 23, 2022). "Dispute arises over P-22's remains as Indigenous people fight for Griffith Park burial". Los Angeles Times. Retrieved December 24, 2022.
56. "Native tribes call for famed mountain lion P-22 to be buried with ceremony near Griffith Park". ABC7 Los Angeles. December 24, 2022. Retrieved December 24, 2022.
57. Sternfield, Marc (December 24, 2022). "Tribal concerns complicate plans for P-22; Natural History Museum says the cougar will not go on display". KTLA. Retrieved December 25, 2022.
58. Dazio, Stefanie (February 4, 2023). "Tribes, researchers debate final fate of P-22, famed LA puma". AP News. Retrieved February 12, 2023.
59. Arce, Ernesto; Ogilvie, Jessica P. (March 6, 2023). "P-22 Has Been Laid To Rest In The Santa Monica Mountains In A Private Ceremony". LAist. Retrieved March 7, 2023.
60. Sternfield, Marc (March 6, 2023). "Famed mountain lion P-22 buried in tribal ceremony". KTLA. Retrieved March 6, 2023.
61. "Famed mountain lion P-22 buried in the Santa Monica Mountains". ABC 7 Los Angeles. March 6, 2023. Retrieved March 6, 2023.
62. "Governor Newsom Statement on Mountain Lion P-22". gov.ca.gov (Press release). Sacramento, California. Office of Governor of the State of California. December 17, 2022.
63. McDaniel, Justine (December 17, 2022). "Iconic L.A. mountain lion euthanized after 'extraordinary life'". The Washington Post.
64. Rainey, James (December 17, 2022). "Mourning 'L.A.'s coolest cat' and celebrating how P-22 changed our relationship with nature". Los Angeles Times.
65. Dahir, Ikran (December 17, 2022). "This Moving Eulogy Of P-22, The Famous Big Cat That Stole Los Angeles's Heart, Is Making Us Tear Up". Buzzfeed News.
66. "Congressman Schiff on the Passing of P-22" (Press release). Office of Congressman Adam Schiff CA 28. December 17, 2022. Retrieved December 21, 2022.
67. Toohey, Grace (December 28, 2022). "Celebration of life for P-22 set for Feb. 4 in Griffith Park". Los Angeles Times. Retrieved January 1, 2023.
68. Schreiner, Casey (October 20, 2022). "L.A.'s most famous mountain lion gets a party this weekend". Los Angeles Times. Retrieved October 28, 2022.
69. "P-22". Natural History Museum.
70. Kevin Andrew Dolak (December 22, 2022). "I Am the Man Behind the Anonymous P-22 Twitter Account". Los Angeles Magazine. Retrieved December 26, 2022.
71. Dickinson, Ian (March 11, 2021). "World's largest wildlife crossing one step closer to becoming a reality". Earth Touch News Network.
72. Von Quednow, Cindy; Riesmeyer, Andy (February 23, 2023). "Los Angeles Public Library releases limited-edition library card honoring P-22". KTLA. Retrieved February 24, 2023.
73. Nelson, Laura J. (February 3, 2023). "A P-22 postage stamp? Schiff kicks off effort to honor L.A.'s celebrity puma". Los Angeles Times. Archived from the original on February 3, 2023. Retrieved February 3, 2023.
74. Cowan, Jill (May 4, 2023). "The Enduring Power of a Big Cat in Star-Obsessed Los Angeles". The New York Times. ISSN 0362-4331. Retrieved May 13, 2023.
75. Guidi, Ruxandra (May 1, 2022). "The lion king of Los Angeles". High Country News. Retrieved May 2, 2022.
76. "Wallis Annenberg Wildlife Crossing". Annenberg Foundation. July 10, 2023.
77. Wagner, Tara Lynn (August 24, 2020). "LA's Most Famous Mountain Lion Stars in New Mural". Spectrum News 1. Archived from the original on November 30, 2021.
78. City News Service (April 19, 2021). "New Watts mural features famed P-22 mountain lion". KNX News 97.1 FM.
79. Keller, Bertram (May 27, 2021). "Mural in Watts Illustrates When Nature Imitates Life". Los Angeles Sentinel.
80. Feraday, Caroline (December 5, 2022). "The P-22 mountain lion is featured in a new mural project in the Conejo Valley". KCLU. Retrieved December 21, 2022.
81. Kurzweil, Tony (October 5, 2022). "Local artist features mountain lion P-22 in Silver Lake mural". KTLA.
82. Scauzillo, Steve (October 4, 2022). "P-22, the 'Hollywood Cat' who lives in Griffith Park, is honored in a mural in Silver Lake". Los Angeles Daily News.
83. Chow, Vivian (December 21, 2022). "'Long Live the King': Mural honoring famed L.A. mountain lion P-22 debuts". KTLA.
84. Tokumatsu, Gordon (October 17, 2024). "Muralist honors legendary mountain lion P-22 with artwork in Hollywood". NBC4.
85. We Heart P-22: A Coloring + Activity Book Celebrating L. A.'s Most Famous Mountain Lion. Narrated Objects. November 2017. ISBN 9780999167007.
86. Medenilla, Klarize (June 2, 2021). "Filipino American-Led 'We Heart L.A. Parks' Celebrates The Rich History Of Los Angeles Parks And The Diverse Communities That Uplift Them". MDWK Magazine. Asian Journal.
87. Guldimann, Suzanne (October 5, 2018). "P-22: The Journey Invites Contemplation about L.A's Big Cats". M'Online. Retrieved October 28, 2022.
88. Lane, Cassandra (November 4, 2020). "READ ABOUT – AND HELP – MOUNTAIN LION P22". L.A. Parent.
89. Zeng, Cady (April 6, 2021). "Environmental Kids' Books for 2021". Publishers Weekly.
90. Westmoore, Jean (February 12, 2021). "Books in Brief: Treasure of the World, The Incredible Painting of Felix Clousseau, Cougar Crossing". The Buffalo News.
91. Schlichenmeyer, Terri (May 24, 2021). "Ring the bell, kids! Summer's here, along with some fantastic reads". The Guam Daily Post.
92. Shively, Kim (July 16, 2016). "The Secret Diary of P-22".
93. The Secret Diary of P-22. Vimeo (Motion picture). 2017.
94. "About". The Cat That Changed America. June 26, 2016. Retrieved October 28, 2022.
95. "S4E6: Stupid Piece of Sh*t". Bojack Horseman. Season 4. Netflix.
96. "Deep Los Angeles Easter Egg". Reddit. September 25, 2017. Archived from the original on December 19, 2022.
    "Bojack Horseman easter eggs". Reddit. September 25, 2017. Archived from the original on December 19, 2022.
97. "This type of wild cat with many names has a superstar in P-22 prowling the canyons of the L.A. area for years". Jeopardy Archive. September 27, 2022. Retrieved December 21, 2022.
98. "This Fool - Sh*t or Get Off the Pot Transcript". TV Show Transcript. Retrieved March 12, 2023.
99. Len, Sophie (December 17, 2022). "LA's Beloved Mountain Lion, P-22, Has Been Euthanized". Secret Los Angeles. Retrieved March 12, 2023.
100. McCarthy, Sean L. (July 28, 2023). "Stream It Or Skip It: 'This Fool' Season 2 On Hulu, Where It's Always Sunny And Foolish In South Central L.A." Decider. Retrieved August 1, 2023.
101. Martin, Dana (November 6, 2018). "Reviews: The Animal Monologues". Stage Raw.
102. Falling James (February 25, 2019). "Amy Raasch Unleashes Her Inner Beast in The Animal Monologues". LA Weekly.
103. Cave, Nick (February 8, 2023). "In light of his death, do you have any lines you'd like to share about this rare, wild creature?". The Red Hand Files. No. 221. Retrieved November 13, 2023.

External links

• P-22's Decade in Griffith Park – Friends of Griffith Park
• Ordeñana, Miguel. "The Sound of Our Griffith Park Mountain Lion: P-22 and the Mysteries of Puma Communication". Los Angeles County Natural History Museum.
• Nijhuis, Michelle (April 20, 2015). "L.A.'s Loneliest Lion". The New Yorker. Retrieved December 13, 2022.