Erechthias is a genus of the fungus mothfamily, Tineidae. Therein, it belongs to the subfamilyErechthiinae, of which it is the type genus. The exact circumscription of this genus is still disputed, but it may encompass more than 150 species.
Systematics and taxonomy
Here, the genus is treated in the wide circumscription (sensu lato) adopted by many authors today, and represents the presumed core group of the Erechthiinae. Delimited thus, Erechthias includes several other genera, some of which have occasionally been treated as independent even by fairly recent authors. They are still rather similar and contain moths that are (at least overwhelmingly) very closely related. Still, they differ in details such as the wing venation – with Erechthiassensu stricto having all veins separate (as opposed to e.g. the Decadarchis group, which has hindwing veins 5 and 6 stalked) – or the clasper's harpe being nude in Erechthias s.str. but bearing a cluster of setae on the costa. However, the female genitals look almost alike in all of them.[1]
Many of these supposedly distinct genera were always considered monotypic and are unlikely to be valid. More notable are Decadarchis (including Caryolestis, Nesoxena, Pantheus, and perhaps others) and Ereunetis (including Lepidobregma and Neodecadarchis); these two are more frequently considered separate genera than other subgroups of Erechthias. The members of the former (sub)genus were in fact at first often placed in Tinea of subfamily Tineinae. Other species of Erechthias were historically assigned to Acridotarsa – also of the Tineinae (E. deloneura) – or to Mesopherna of the Myrmecozelinae (E. epomadia). E. glyphidaula has been a particular source of confusion; even veteran researcher Edward Meyrick, in some of his last works, no less than three times established a new monotypic genus for this species.[2]
More unusually, some species of Erechthias were initially mistaken as members of the cosmet moth genus Cosmopteryx (E. cyanosticta) and the ermine moth genus Argyresthia (E. zebrina); these genera are basalDitrysia not particularly closely related to Erechthias. The enigmatic "genus" Acrocenotes – a single species initially held to belong to the Plutellidae, which are also not close relatives of Erechthias – is also included here in the present treatment.[3]
Finally, the actual delimitation of Erechthias against related genera such as Comodica still needs to be determined. For example, whether the Erechthiinae Callicerastis stagmatias and Mecomodica fullawayi are justifiably separated in their monotypic genera or better included in Erechthias (as are all the other species once placed in Callicerastis) is disputed; the latter in particular seems to be somewhat intermediate between Comodica and Erechthias.[5]
Clarke, John Frederick Gates (1986): Pyralidae and Microlepidoptera of the Marquesas Archipelago. Smithsonian Contributions to Zoology416: 1-485. PDF fulltextArchived 2012-03-27 at the Wayback Machine (214 MB!)
