Corycidia is defined as the least inclusive clade containing Amphizonella, Diplochlamys and Trichosphaerium. It was discovered in 2017 through a phylogenomic study by Senghuo Kang and coauthors, published in the journal Molecular Biology and Evolution.[1] It is supported by independent analyses.[2] As of 2019, it is accepted by the International Society of Protistologists as part of the modern cladistic classification of eukaryotes, although under the alternative spelling Corycida.[3] The name Corycidia is derived from Latin corium 'leathery', referring to the "leathery" test that members of the clade have.[1]
A total of 7 genera are included within Corycidia, traditionally organized into two families: Trichosphaeriidae (one genus Trichosphaerium), under the monotypic order Trichosida, and Microcoryciidae (six genera), previously assigned to the order Arcellinida. Only three genera are firmly placed in Corycidia through phylogenetic analyses: Amphizonella, Diplochlamys and Trichosphaerium.[1] The position of the genera Penardochlamys, Microcorycia, Zonomyxa and Parmulina, listed as part of the family Microcoryciidae,[4] is not clear. They have been tentatively placed in Corycidia due to a morphological resemblance, but it has not been demonstrated with molecular data.[3]
^ abcdefKang, Seungho; Tice, Alexander K; Spiegel, Frederick W; Silberman, Jeffrey D; Pánek, Tomáš; Čepička, Ivan; Kostka, Martin; Kosakyan, Anush; Alcântara, Daniel M C; Roger, Andrew J; Shadwick, Lora L; Smirnov, Alexey; Kudryavtsev, Alexander; Lahr, Daniel J G; Brown, Matthew W (September 2017). "Between a Pod and a Hard Test: The Deep Evolution of Amoebae". Molecular Biology and Evolution. 34 (9): 2258–2270. doi:10.1093/molbev/msx162. PMC5850466. PMID28505375.
^ abAdl SM, Bass D, Lane CE, Lukeš J, Schoch CL, Smirnov A, Agatha S, Berney C, Brown MW, Burki F, Cárdenas P, Čepička I, Chistyakova L, del Campo J, Dunthorn M, Edvardsen B, Eglit Y, Guillou L, Hampl V, Heiss AA, Hoppenrath M, James TY, Karnkowska A, Karpov S, Kim E, Kolisko M, Kudryavtsev A, Lahr DJG, Lara E, Le Gall L, Lynn DH, Mann DG, Massana R, Mitchell EAD, Morrow C, Park JS, Pawlowski JW, Powell MJ, Richter DJ, Rueckert S, Shadwick L, Shimano S, Spiegel FW, Torruella G, Youssef N, Zlatogursky V, Zhang Q (2019). "Revisions to the Classification, Nomenclature, and Diversity of Eukaryotes". Journal of Eukaryotic Microbiology. 66 (1): 4–119. doi:10.1111/jeu.12691. PMC6492006. PMID30257078.
^Kosakyan, Anush; Gomaa, Fatma; Lara, Enrique; J.G. Lahr, Daniel (2016). "Current and future perspectives on the systematics, taxonomy and nomenclature of testate amoebae". European Journal of Protistology. 55 (Part B): 105–117. doi:10.1016/j.ejop.2016.02.001.