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Alan Grafen
Alan Grafen FRS is a Scottish ethologist and evolutionary biologist. He currently teaches and undertakes research at St John's College, Oxford.[1] Along with regular contributions to scientific journals, Grafen is known publicly for his work as co-editor (with Mark Ridley) of the 2006 festschrift Richard Dawkins: How a Scientist Changed the Way We Think,[2] honouring the achievements of his colleague and former academic advisor. He has worked extensively in the field of biological game theory, and, in 1990, devised a model showing that Zahavi's well-known handicap principle could theoretically exist in natural populations.[3][4] He also published a seminal paper in the field of phylogenetic comparative methods, in which he demonstrated how the tools of generalized least squares could be applied to perform phylogenetically informed statistical analyses.[5] Grafen was elected a Fellow of the Royal Society in 2011.[6] Bibliography
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