Subfamily of true bugs
Pentatominae is a subfamily of Pentatomidae , a family of shield bugs. This subfamily is the largest one within the Pentatomidae, having 4937 species classified in 938 genera.[ 1] Species in this subfamily are phytophages and several of them are considered agricultural pests.[ 2] Some invasive pentatomines such as Halyomorpha halys and Bagrada hilaris have been considered household pests.[ 3] Higher systematics of the group have been revised by Rider et al.[ 4]
Tribes and genera
Tribes within this subfamily include:[ 5]
Aelini Douglas & Scott, 1865
Aelia Fabricius, 1803
Aeliopsis Bergevin, 1931
Neottiglossa Kirby, 1837
Aeliosoma Baehr, 1985
Aeptus Dallas, 1851
Caridillus Bergroth, 1909
Eribotes (bug) Stål, 1867
Hillieria Distant, 1910
Mariomella Faúndez & Rider, 2019
Menestheus (bug) Stål, 1867
Paramenestheus Breddin, 1900
Pseudaelia Distant, 1910
Aeschrocorini Distant, 1902
Aeschrocoris Bergroth, 1887
Aeschrus Spinola, 1850
Chraesus Cachan, 1952
Geomorpha (bug) Bergroth, 1893
Risbecella Schouteden, 1942
Scylax (bug) Distant, 1887
Tyoma Spinola, 1850
Tyomana Miller, 1952
Agaeus Dallas, 1851
Agonoscelidini Atkinson, 1888
Agonoscelis Spinola, 1837
Ambohicorypha Cachan, 1952
Belopis Distant, 1879
Bolaca Walker, 1867
Halyabbas Distant, 1900
Madecorypha Cachan, 1952
Sennertus Distant, 1900
Antestiini Distant, 1902 - selected genera:
Aulacetrini Mulsant & Rey, 1866
Holcogaster Fieber, 1860
Axiagastini Atkinson, 1888
Acesines Stål, 1876
Axiagastus Dallas, 1851
Indrapura (bug) China, 1928
Oncotropis Stål, 1871
Bathycoelia Amyot & Serville, 1843
Adelolcus Bergroth, 1893
Benia (bug) Schouteden, 1916
Boerias Kirkaldy, 1909
Cappaea Ellenrieder, 1862
Caura (bug) Stål, 1865
Cauromorpha Jeannel, 1913
Erlangerella Schouteden, 1905
Halycorypha Jeannel, 1913
Halydicoris Jeannel, 1913
Halyoides Cachan, 1952
Halyomorpha Mayr, 1864
Homalogonia Jakovlev, 1876
Hymenomaga Karsch, 1892
Lerida (bug) Karsch, 1894
Leridella Jeannel, 1913
Lokaia Linnavuori, 1975
Mabusana Distant, 1912
Massocephalus Dallas, 1851
Paralerida Linnavuori, 1982
Prytanicoris Gross, 1978
Tolumnia Stål, 1867
Tripanda Berg, 1899
Tropicorypha Mayr, 1864
Veterna Stål, 1865
Carpocorini Mulsant & Rey, 1866 selected genera:
Antheminia Mulsant & Rey, 1866
Carpocoris Kolenati, 1846
Chlorochroa Stål, 1872
Dolycoris Mulsant & Rey, 1866
Graphorn Faúndez, Rider, & Carvajal, 2017
Peribalus Mulsant & Rey, 1866
Rubiconia Dohrn, 1860
Thestral Faúndez & Rider, 2014
Catacanthini Atkinson, 1888
Arocera Spinola, 1837
Boea (bug) Walker, 1867
Catacanthus Spinola, 1837
Chalcocoris Dallas, 1851
Rhyssocephala Rider, 1992
Runibia Stål, 1861
Vitruvius (bug) Distant, 1901
Vulsirea Spinola, 1837
Caystrini Ahmad & Afzal, 1979
Caystrus Stål, 1861
Cephaloplatus White, 1842
Heissocoris Ahmad & Afzal, 1979
Hippotiscus Bergroth, 1906
Kyrtalus Van Duzee, 1929
Lyalocoris Zahid, 2006
Mcdonaldiana Zahid, 2006
Neodius (Stål, 1867)
Neoparodius Zahid, 2006
Neoriazocoris Zahid, 2006
Paramecocoris Stål, 1853
Parodius (bug) Distant, 1900
Praetextatus (bug) Distant, 1901
Riazocoris Ahmad & Afzal, 1979
Valescus Distant, 1901
Chlorocorini Rider, Greve, Schwertner & Grazia, 2018
Arvelius Spinola, 1837
Chlorocoris Spinola, 1837
Chloropepla Stål, 1867
Eludocoris Thomas, 1992
Fecelia Stål, 1872
Mayrinia Horváth, 1925
Rhyncholepta Bergroth, 1911
Coquereliini Cachan, 1952
Cleoqueria Cachan, 1952
Coquerelia Signoret, 1861
Coquerelidea Reuter, 1887
Neocoquerelidea Cachan, 1952
Degonetini Azim & Shafee, 1984
Degonetus Distant, 1902
Diemeniini Kirkaldy, 1909 - selected genera:
Diplostirini Distant, 1902
Ambiorix (bug) Stål, 1876
Diplostira Dallas, 1851
Diploxyini Atkinson, 1888
Acoloba Spinola, 1850
Aeladria Cachan, 1952
Diploxys Amyot & Serville, 1843
Eurysaspidini Atkinson, 1888
Eurysaspis Signoret, 1851
Flaminia (bug) Stål, 1865
Eysarcorini Mulsant & Rey, 1866
Adria (bug) Stål, 1876
Aspavia Stål, 1865
Cachanocoris Rider, 1998
Carbula Stål, 1864
Corisseura Cachan, 1952
Cratonotus Distant, 1879
Durmia Stål, 1865
Dymantiscus Hsiao, 1981
Empiesta Linnavuori, 1982
Eysarcoris Hahn, 1834
Hermolaus Distant, 1902
Mulungua Linnavuori, 1982
Pseudolerida Schouteden, 1958
Sepontia Stål, 1865
Sepontiella Miyamoto, 1990
Spermatodes Bergroth, 1914
Stagonomus Gorski, 1852
Halyini Amyot & Serville, 1843 - selected genera:
Hoplistoderini Atkinson, 1888
Alcimocoris Bergroth, 1891
Glottaspis Bergroth, 1918
Hoplistodera Westwood, 1837
Paracritheus Bergroth, 1891
Paralcimocoris Zhang & Lin, 1988
Stachyomia Stål, 1871
Taurodes Dallas, 1851
Lestonocorini Ahmad & Mohammad, 1980
Dardjilingia Yang, 1936
Gynenica Dallas, 1851
Lestonocoris Ahmad & Mohammad, 1980
Neogynenica Yang, 1935
Umgababa (bug) Leston, 1953
Mecidea Dallas, 1851
Memmia Stål, 1865
Actuarius (bug) Distant, 1901
Aegaleus (bug) Stål, 1876
Amphimachus (bug) Stål, 1867
Anchesmus Stål, 1876
Apines Dallas, 1851
Aspideurus Signoret, 1880
Banya (bug) Schouteden, 1916
Brachycoris Stål, 1871
Carvalhocoris Leston, 1953
Cresphontes (bug) Stål, 1867
Dabessus Distant, 1902
Decellella Schouteden, 1964
Desertomenida Kiritshenko, 1914
Dunnius Distant, 1902
Elanela Rolston, 1980
Eurymenida Bergroth, 1917
Gwea Schouteden, 1958
Keleacoris Rider & Rolston, 1995
Lathraedoeus Breddin, 1900
Menida Motschulsky, 1861
Neostrachia Saunders, 1877
Phavorinus Distant, 1902
Rio (bug) Kirkaldy, 1909
Saceseurus Breddin, 1900
Sciomenida Gross, 1976
Solenocoris Oshanin, 1922
Udonga Distant, 1921
Aednulus Breddin, 1901
Aednus Dallas, 1851
Arniscus Distant, 1899
Delegorguella Spinola, 1850
Dictyotus Dallas, 1851
Dictytotus Dallas, 1851
Dissocolpus Bergroth, 1906
Dollingiana Ahmad & Kamaluddin, 1986
Dymantis Stål, 1861
Ennius (bug) Stål, 1861
Erachtheus Stål, 1861
Humria Linnavuori, 1975
Jitka (bug) Kment, 2015
Laprius Stål, 1861
Lodosia Ahmad & Önder, 1996
Munshiana Ahmad & Kamaluddin, 1987
Myrochea Amyot & Serville, 1843
Neococalus Bergroth, 1891
Neodorpius Ahmad & Afzal, 1989
Paradictyotus Gross, 1975
Pretorius (bug) Distant, 1898
Stysicoris Ahmad & Kamaluddin, 1985
Tholosanus Distant, 1899
Utheria Gross, 1975
Nealeria Bergroth, 1893
Paraleria Reuter, 1887
Nezarini Atkinson, 1888 - selected genera:
Opsitomini Cachan, 1952
Opsitoma Cachan, 1952
Pentamyrmexini Rider & Brailovsky, 2014 (monotypic, Thailand)
Pentatomini Leach, 1815 - selected genera:
Phricodini Cachan, 1952
Phricodus Spinola, 1839
Piezodorini Atkinson, 1888
Anaximenes (bug) Stål, 1876
Chaubattiana Distant, 1912
Pausias (bug) Jakovlev, 1905
Piezodorus Fieber, 1860
Procleticini Pennington, 1920
Aleixus McDonald, 1981
Brepholoxa Van Duzee, 1904
Dendrocoris Bergroth, 1891
Lobepomis Berg, 1891
Neoderoploa Pennington, 1922
Odmalea Bergroth, 1914
Parodmalea Rider, 1994
Procleticus Berg, 1891
Pseudobebaeus Distant, 1911
Terania Pirán, 1963
Thoreyella Spinola, 1850
Araducta Walker, 1867
Avicenna (bug) Distant, 1901
Biprorulus Breddin, 1900
Cuspicona Dallas, 1851
Diaphyta Bergroth, 1891
Everardia (bug) Gross, 1975
Hoffmanseggiella Spinola, 1850
Koogobatha Distant, 1910
Morna (bug) Stål, 1867
Mygoodano Distant, 1910
Ocirrhoe Stål, 1876
Parocirrhoe Gross, 1975
Pegala Stål, 1867
Petalaspis Bergroth, 1916
Pugione Stål, 1867
Rhynchocoris Westwood, 1837
Utana Distant, 1900
Vitellus Stål, 1865
Rolstoniellini Rider, 1997
Agathocles (bug) Stål, 1876
Amasenus Stål, 1863
Amatembuna Distant, 1910
Critheus Stål, 1867
Exithemus Distant, 1902
Rolstoniellus Rider, 1997
Sciocorini Amyot & Serville, 1843 selected genera:
Strachiini Mulsant & Rey, 1866 selected genera:
Triplatyxini Cachan, 1952
Anoano Cachan, 1952
Nene (bug) Kment, 2015
Tricompastes Cachan, 1952
Triplatyx Horváth, 1904
Amphidexius Bergroth, 1918
Bachesua Gross, 1975
Birna McDonald, 2006
Brizocoris Gross, 1975
Buthumka Gross, 1975
Cooperocoris Gross, 1975
Dysnoetus Bergroth, 1918
Eremophilacoris Gross, 1975
Gadarscama Reuter, 1887
Ippatha Distant, 1910
Kitsonia Gross, 1975
Kitsoniocoris Rider, 1998
Kumbutha Distant, 1910
Lakhonia Yang, 1936
Macrocarenoides Gross, 1975
Macrocarenus Stål, 1873
Mcphersonarcys Thomas, 2012
Muscanda Walker, 1868
Ochisme Kirkaldy, 1904
Ooldeon Gross, 1975
Patanius Rolston, 1987
Pirricoris Gross, 1975
Poecilotoma Dallas, 1851
Riaziana Hasan, Afzal & Ahmad, 1989
Senectius Rolston, 1987
Tepperocoris Gross, 1975
Thryptomenecoris Gross, 1975
Trachyops Dallas, 1851
Turrubulana Distant, 1910
Xiengia Distant, 1921
Zouicoris Zheng, 1986
Notable species
References
^ Faúndez, E. I.; Rider, D. A.; Garcete-Barret, B. (2018). "New records of Alcaeorrhynchus grandis (Dallas, 1851)(Heteroptera: Pentatomidae) from Paraguay, with description of a teratological case" . Boletín del Museo Nacional de Historia Natural del Paraguay . 22 (1): 27– 29.
^ Faúndez, E. I.; Rider, D. A. (2017). "The brown marmorated stink bug Halyomorpha halys (Stål, 1855)(Heteroptera: Pentatomidae) in Chile" . Arquivos Entomolóxicos . 17 : 305– 307.
^ Faúndez, E. I. (2018). "From agricultural to household pest: The case of the painted bug Bagrada hilaris (Burmeister)(Heteroptera: Pentatomidae) in Chile". Journal of Medical Entomology . 55 (5): 1365– 1368. doi :10.1093/jme/tjy091 . PMID 29931070 .
^ Rider, D.A., Schwertner, C.F., Vilimová, J., Redei, D., Kment, P. & D.B. Thomas. 2018. Pp. 25-204. Higher systematics of Pentatomidea in McPherson, J. E. (Ed.). Invasive Stink Bugs and Related Species (Pentatomoidea): Biology, Higher Systematics, Semiochemistry, and Management. CRC Press, Florida.
^ Zicha, Ondrej. "BioLib: Biological library" . www.biolib.cz .
External links