en Cereinae

Cereinae
	Cereus repandus
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Order:	Caryophyllales
Family:	Cactaceae
Subfamily:	Cactoideae
Tribe:	Cereeae
Subtribe:	Cereinae; Britton & Rose
Type genus
	Cereus;
Genera
	See text.

Cereinae is a subtribe of cacti in the tribe Cereeae, subfamily Cactoideae. It is one of the six subtribes into which the tribe Cereeae was divided in 2023, the others being the monotypic Aylosterinae, Gymnocalyciinae, and Uebelmanniinae, along with Rebutiinae and Trichocereinae.^[1]

Genera

14 genera were firmly included in subtribe Cereinae in a 2021 study.^[2] This included Stephanocereus, treated as a synonym of Arrojadoa by Plants of the World Online as of September 2023^[update].^[3] Espostoopsis was not included in the 2021 study, but was noted to have been placed in Cereinae in many previous studies, including that by Lendel in 2013.^[2]^[4] Serrulatocereus was also not included in the 2021 study. It may not be distinct from Cereus as molecular phylogenetic studies have placed it in that genus, a placement accepted by some sources.^[5] A 2023 molecular phylogenetic study found that Stetsonia, formerly placed in subtribe Rebutiinae, belonged in Cereinae.^[1]

Image	Genus	Living species
	Arrojadoa Britton & Rose, (1920) (including Stephanocereus)	Arrojadoa albiflora Arrojadoa bahiensis Arrojadoa dinae Arrojadoa eriocaulis Buining & Brederoo Arrojadoa leucostele (Gürke) Anceschi & Magli Arrojadoa marylanae Arrojadoa multiflora F.Ritter Arrojadoa olsthoorniana Hofacker, M.Machado & R.Pontes Arrojadoa penicillata Arrojadoa rhodantha
	Brasilicereus Backeb.	Brasilicereus estevesii Brasilicereus markgrafii Brasilicereus phaeacanthus
	Cereus Mill.	25 species: Cereus aethiops Haw. Cereus bicolor Rizzini & A.Mattos Cereus fernambucensis Lem. Cereus forbesii C.F.Först. Cereus fricii Backeb. Cereus hexagonus (L.) Mill. Cereus hildmannianus K.Schum. Cereus horrispinus Backeb. Cereus insularis Hemsl. Cereus jamacaru DC. Cereus lamprospermus K.Schum. Cereus lanosus (F.Ritter) P.J.Braun Cereus lepidotus Salm-Dyck Cereus mortensenii (Croizat) D.R.Hunt & N.P.Taylor Cereus pachyrrhizus K.Schum. Cereus phatnospermus K.Schum. Cereus pierre-braunianus Esteves Cereus repandus (L.) Mill. Cereus saddianus (Rizzini & A.Mattos) P.J.Braun Cereus spegazzinii F.A.C.Weber Cereus stenogonus K.Schum. Cereus trigonodendron K.Schum. ex Vaupel Cereus vargasianus Cárdenas Cereus yungasensis A.Fuentes & Quispe
	Cipocereus F.Ritter	Cipocereus bradei Cipocereus crassisepalus Cipocereus laniflorus Cipocereus minensis Cipocereus pusilliflorus
	Coleocephalocereus Backeb.	Coleocephalocereus aureus F.Ritter Coleocephalocereus buxbaumianus Buining Coleocephalocereus buxbaumianus subsp. buxbaumianus. Coleocephalocereus buxbaumianus subsp. flavisetus (F.Ritter) N.P.Taylor & Zappi Coleocephalocereus fluminensis (Miq.) Backeb. Coleocephalocereus fluminensis subsp. fluminensis Coleocephalocereus fluminensis subsp. decumbens (F.Ritter) N.P.Taylor & Zappi Coleocephalocereus goebelianus (Vaupel) Buining Coleocephalocereus pluricostatus Buining & Brederoo Coleocephalocereus purpureus (Buining & Brederoo) F.Ritter
	Discocactus Pfeiff. 1837	Discocactus bahiensis Britton & Rose Discocactus boliviensis Backeb. ex Buining Discocactus buenekeri W.R.Abraham Discocactus catingicola Buining & Brederoo Discocactus diersianus Esteves Pereira Discocactus fariae-peresii P.J.Braun Discocactus ferricola Buining & Brederoo Discocactus hartmannii (K.Schum.) Britton & Rose Discocactus heptacanthus (Barb.Rodr.) Britton & Rose Discocactus horstii Buining & Brederoo Discocactus petr-halfarii Zachar Discocactus placentiformis (Lehm.) K.Schum. Discocactus pseudoinsignis N.P.Taylor & Zappi Discocactus zehntneri Britton & Rose
	Espostoopsis Buxb. 1968	Espostoopsis dybowskii (Rol.-Goss.) Buxb.
	Facheiroa Britton & Rose 1920	Facheiroa braunii Esteves Facheiroa cephaliomelana Buining & Brederoo Facheiroa squamosa (Gürke) P.J.Braun & Esteves Facheiroa ulei (K.Schum.) Werderm.
	Leocereus Britton & Rose 1920	Leocereus bahiensis Britton & Rose
	Melocactus Link & Otto	52 species: Melocactus acunae León Melocactus × albicephalus Buining & Brederoo Melocactus andinus R.Gruber ex N.P.Taylor Melocactus azureus Buining & Brederoo Melocactus bahiensis (Britton & Rose) Luetzelb. Melocactus bellavistensis Rauh & Backeb. Melocactus braunii Esteves Melocactus brederooanus Buining Melocactus broadwayi (Britton & Rose) Houghton Melocactus caroli-linnaei N.P.Taylor Melocactus conoideus Buining & Brederoo Melocactus curvispinus Pfeiff. Melocactus deinacanthus Buining & Brederoo Melocactus ernestii Vaupel Melocactus estevesii P.J.Braun Melocactus evae Z.Mészáros Melocactus ferreophilus Buining & Brederoo Melocactus glaucescens Buining & Brederoo Melocactus harlowii (Britton & Rose) Vaupel Melocactus heimenii P.J.Braun & Gonç.Brito Melocactus holguinensis Areces Melocactus × horridus Werderm. Melocactus inconcinnus Buining & Brederoo Melocactus intortus (Mill.) Urb. Melocactus lagunaensis (Z.Mészáros) D.Barrios & Majure Melocactus lanssensianus P.J.Braun Melocactus lemairei (Monv. ex Lem.) Miq. ex Lem. Melocactus levitestatus Buining & Brederoo Melocactus macracanthos (Salm-Dyck) Link & Otto Melocactus matanzanus León Melocactus mazelianus Říha Melocactus nagyi Z.Mészáros Melocactus neoviridescens Guiggi Melocactus neryi K.Schum. Melocactus oreas Miq. Melocactus pachyacanthus Buining & Brederoo Melocactus paucispinus Heimen & R.J.Paul Melocactus pedernalensis M.M.Mejía & R.G.García Melocactus perezassoi Areces Melocactus peruvianus Vaupel Melocactus praerupticola Areces Melocactus pruinosus Werderm. Melocactus radoczii Z.Mészáros Melocactus salvadorensis Werderm. Melocactus santiagoensis D.Barrios & Majure Melocactus schatzlii H.Till & R.Gruber Melocactus sergipensis N.P.Taylor & Meiado Melocactus smithii (Alexander) Buining ex G.D.Rowley Melocactus stramineus Suringar Melocactus × trachycephalus Suringar Melocactus violaceus Pfeiff. Melocactus zehntneri (Britton & Rose) Luetzelb.
	Micranthocereus Backeb.	Micranthocereus albicephalus (Buining & Brederoo) F.Ritter Micranthocereus auri-azureus Buining & Brederoo Micranthocereus dolichospermaticus (Buining & Brederoo) F.Ritter Micranthocereus estevesii (Buining & Brederoo) F.Ritter Micranthocereus flaviflorus Buining & Brederoo Micranthocereus hofackerianus (P.J.Braun & Esteves) M.Machado Micranthocereus polyanthus (Werderm.) Backeb. Micranthocereus polyanthus subsp. alvinii M.Machado & Hofacker Micranthocereus polyanthus subsp. polyanthus Micranthocereus purpureus (Gürke) F.Ritter Micranthocereus streckeri Van Heek & Van Criek. Micranthocereus violaciflorus Buining
	Pilosocereus Byles & G. D. Rowley	51 species: Pilosocereus albisummus P.J.Braun & Esteves Pilosocereus alensis (F.A.C.Weber) Byles & G.D.Rowley Pilosocereus armatus (Otto ex Pfeiff.) A.R.Franck Pilosocereus arrabidae (Lem.) Byles & G.D.Rowley Pilosocereus aureispinus (Buining & Brederoo) F.Ritter Pilosocereus aurilanatus F.Ritter Pilosocereus aurisetus (Werderm.) Byles & G.D.Rowley Pilosocereus azulensis N.P.Taylor & Zappi Pilosocereus bohlei Hofacker Pilosocereus brasiliensis (Britton & Rose) Backeb. Pilosocereus brooksianus (Britton & Rose) Byles & G.D.Rowley Pilosocereus catingicola (Gürke) Byles & G.D.Rowley Pilosocereus chrysacanthus (F.A.C.Weber ex K.Schum.) Byles & G.D.Rowley Pilosocereus chrysostele (Vaupel) Byles & G.D.Rowley Pilosocereus collinsii (Britton & Rose) Byles & G.D.Rowley Pilosocereus colombianus (Rose) Byles & G.D.Rowley Pilosocereus curtisii (Otto) A.R.Franck Pilosocereus densiareolatus F.Ritter Pilosocereus diersianus (Esteves) P.J.Braun Pilosocereus flavipulvinatus (Buining & Brederoo) F.Ritter Pilosocereus flexibilispinus P.J.Braun & Esteves Pilosocereus floccosus Byles & G.D.Rowley Pilosocereus fulvilanatus (Buining & Brederoo) F.Ritter Pilosocereus gaumeri (Britton & Rose) Backeb. Pilosocereus glaucochrous (Werderm.) Byles & G.D.Rowley Pilosocereus jamaicensis Proctor ex A.R.Franck & al. Pilosocereus kanukuensis (Alexander) Leuenb. Pilosocereus lanuginosus (L.) Byles & G.D.Rowley Pilosocereus leucocephalus (Poselg.) Byles & G.D.Rowley Pilosocereus machrisii (E.Y.Dawson) Backeb. Pilosocereus magnificus (Buining & Brederoo) F.Ritter Pilosocereus millspaughii (Britton) Byles & G.D.Rowley Pilosocereus mollispinus P.J.Braun & Esteves Pilosocereus moritzianus (Otto) Byles & G.D.Rowley Pilosocereus multicostatus F.Ritter Pilosocereus oligolepis (Vaupel) Byles & G.D.Rowley Pilosocereus pachycladus F.Ritter Pilosocereus parvus (Diers & Esteves) P.J.Braun Pilosocereus pentaedrophorus (Labour.) Byles & G.D.Rowley Pilosocereus piauhyensis (Gürke) Byles & G.D.Rowley Pilosocereus polygonus (Lam.) Byles & G.D.Rowley Pilosocereus purpusii (Britton & Rose) Byles & G.D.Rowley Pilosocereus pusillibaccatus P.J.Braun & Esteves Pilosocereus quadricentralis (E.Y.Dawson) Backeb. Pilosocereus robinii (Lem.) Byles & G.D.Rowley Pilosocereus splendidus F.Ritter Pilosocereus ×subsimilis Rizzini & A.Mattos Pilosocereus tillianus R.Gruber & Schatzl Pilosocereus ulei (K.Schum.) Byles & G.D.Rowley Pilosocereus vilaboensis (Diers & Esteves) P.J.Braun Pilosocereus zappiae Lavor & Calvente
	Praecereus Buxb.	Praecereus euchlorus Praecereus saxicola
	Serrulatocereus Guiggi	Serrulatocereus serruliflorus
	Stetsonia Britt. & Rose	Stetsonia coryne
	Xiquexique Lavor, Calvente & Versieux 2020	Xiquexique frewenii (Zappi & N.P.Taylor) Lavor & Calvente Xiquexique gounellei (F.A.C.Weber ex K.Schum.) Lavor & Calvente Xiquexique × heptagonus N.P.Taylor & Albuq.-Lima Xiquexique tuberculatus (Werderm.) Lavor & Calvente

References

^ ^a ^b Romeiro-Brito, Monique; Taylor, Nigel P.; Zappi, Daniela C.; Telhe, Milena C.; Franco, Fernando F. & Moraes, Evandro M. (2023). "Unravelling phylogenetic relationships of the tribe Cereeae using target enrichment sequencing". Annals of Botany. 132 (5): 989–1006. doi:10.1093/aob/mcad153.
^ ^a ^b Fantinati, Mariana R.; Soffiatti, Patricia & Calvente, Alice (2021), "A New Phylogenetic Hypothesis for Cereinae (Cactaceae) Points to a Monophyletic Subtribe" (PDF), Systematic Botany, 46 (3): 689–699, doi:10.1600/036364421X16312068417020, retrieved 2023-09-28
^ "Arrojadoa Britton & Rose", Plants of the World Online, Royal Botanic Gardens, Kew, retrieved 2023-09-29
^ Lendel, Anita (2013). South American Cacti in time and space: studies on the diversification of the tribe Cereeae, with particular focus on subtribe Trichocereinae (Cactaceae) (Dr. sc. nat. thesis). University of Zurich. doi:10.5167/UZH-93287. Retrieved 2023-10-10.
^ Hoxey, Paul; Gdaniec, Andrew; Hernández, José Dolores; Guzmán, Mario; Corniel, Wilmer & Culham, Alastair (2023). "Illustrated Checklist of the Cacti of Hispaniola (Haiti and the Dominican Republic) with Keys to Genera and Species". Caribbean Journal of Science. 53 (2): 85–197. doi:10.18475/cjos.v53i2.a3.

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[RomeTaylZappTelh23-1] Romeiro-Brito, Monique; Taylor, Nigel P.; Zappi, Daniela C.; Telhe, Milena C.; Franco, Fernando F. & Moraes, Evandro M. (2023). "Unravelling phylogenetic relationships of the tribe Cereeae using target enrichment sequencing". Annals of Botany. 132 (5): 989–1006. doi:10.1093/aob/mcad153.

[FantSoffCalv21-2] Fantinati, Mariana R.; Soffiatti, Patricia & Calvente, Alice (2021), "A New Phylogenetic Hypothesis for Cereinae (Cactaceae) Points to a Monophyletic Subtribe" (PDF), Systematic Botany, 46 (3): 689–699, doi:10.1600/036364421X16312068417020, retrieved 2023-09-28

[POWO_297007-2-3] "Arrojadoa Britton & Rose", Plants of the World Online, Royal Botanic Gardens, Kew, retrieved 2023-09-29

[Lendel2013-4] Lendel, Anita (2013). South American Cacti in time and space: studies on the diversification of the tribe Cereeae, with particular focus on subtribe Trichocereinae (Cactaceae) (Dr. sc. nat. thesis). University of Zurich. doi:10.5167/UZH-93287. Retrieved 2023-10-10.

[HoxeGdanHernGuzm23-5] Hoxey, Paul; Gdaniec, Andrew; Hernández, José Dolores; Guzmán, Mario; Corniel, Wilmer & Culham, Alastair (2023). "Illustrated Checklist of the Cacti of Hispaniola (Haiti and the Dominican Republic) with Keys to Genera and Species". Caribbean Journal of Science. 53 (2): 85–197. doi:10.18475/cjos.v53i2.a3.

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