2012 in arthropod paleontology
Overview of the events of 2012 in arthropod paleontology
List of years in arthropod paleontology
This list of fossil arthropods described in 2012 is a list of new taxa of trilobites , fossil insects , crustaceans , arachnids and other fossil arthropods of every kind that have been described during the year 2012. The list only includes taxa at the level of genus or species .
Arachnids
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Ampezzoa [ 1]
Gen. et sp. nov.
Valid
Lindquist & Grimaldi in Schmidt et al.
Carnian
Italy
A mite . The type species is Ampezzoa triassica .
Archaelagonops [ 2]
Gen. et sp. nov
Valid
Wunderlich
Cretaceous
Burmese amber
Myanmar
A member of Lagonomegopidae . Type species is A. salticoides .
Baltleucauge propinqua [ 3]
Sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
A long-jawed orb weaver
Craspedisia yapchoontecki [ 4]
Sp. nov.
Valid
Penney et al.
Miocene
Dominican amber
Dominican Republic
A theridiid
Eodoter lonimammillae [ 3]
Sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
A sac spider
Eohalinobius calefactus [ 3]
Sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
A succinomid
Eohalinobius hiddenseeensis [ 3]
Sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
A succinomid
Eohalinobius patina [ 3]
Sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
A succinomid
?Eophantes seorsum [ 3]
Sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
A linyphiid
Ephalmator tredecim [ 3]
Sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
An ephalmatorid
Ero (Succinero) clunis [ 3]
Sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
A pirate spider
Ero (Succinero) gracilitibialis [ 3]
Sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
A pirate spider
Ero (Succinero) veta [ 3]
Sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
A pirate spider
Geratonephila [ 5]
Gen. et sp. nov.
jr synonym
Poinar in Poinar & Buckley
Early Cretaceous
Burmese amber
Myanmar
A nephiline . The type species is G. burmanica . Moved to Nephila in 2015 .
Nephila burmanica
Graea magnocoli [ 3]
Sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
An orb-weaver spider
Hypotheridiosoma [ 2]
Gen. et sp. nov
Valid
Wunderlich
Cretaceous
Burmese amber
Myanmar
Originally classified as a ray spider , but subsequently transferred to the araneoid family Zarqaraneidae .[ 6] The type species is H. paracymbium .
Korearachne [ 7]
gen et sp nov
Valid
Selden et al. .
Early Cretaceous
Jinju Formation
South Korea
A member of Araneomorphae , possibly a lycosoid . The type species is K. jinju .
Lagonoburmops [ 2]
Gen. et sp. nov
Valid
Wunderlich
Cretaceous
Burmese amber
Myanmar
A member of Lagonomegopidae . The type species is L. plumosus .
Leclercera longissipes [ 2]
Sp. nov
Jr synonym
Wunderlich
Cretaceous
Burmese amber
Myanmar
A member of Psilodercidae . Originally described as a species of Leclercera , but subsequently transferred to the genus Priscaleclercera .[ 8]
Leclercera spicula [ 2]
Sp. nov
Jr synonym
Wunderlich
Cretaceous
Burmese amber
Myanmar
A member of Psilodercidae . Originally described as a species of Leclercera , but subsequently transferred to the genus Priscaleclercera .[ 8]
Leviunguis [ 2]
Gen. et sp. nov
Valid
Wunderlich
Cretaceous
Burmese amber
Myanmar
Originally classified as a ray spider , subsequently transferred to the separate araneoid family Leviunguidae .[ 6] The type species is L. bruckschi .
Mesobunus dunlopi [ 9]
Sp. nov.
Valid
Giribet et al.
Middle Jurassic
Daohugou Beds
China
A sclerosomatid harvestman .
Myanlagonops [ 2]
Gen. et sp. nov
Valid
Wunderlich
Cretaceous
Burmese amber
Myanmar
A member of Lagonomegopidae . The type species is M. gracilipes .
Ocululoborus [ 2]
Gen. et sp. nov
Valid
Wunderlich
Cretaceous
Burmese amber
Myanmar
A member of Uloboridae . The type species is O. curvatus .
Orchestina (Baltorchestina) angulata [ 10]
Nom. nov
Valid
Wunderlich
Eocene
Bitterfeld amber
Germany
An oonopid ; a replacement name for Orchestina (Baltorchestina) rectangulata Wunderlich (2011).
Orchestina gappi [ 11]
sp nov
Valid
Saupe et al. .
Latest Albian or earliest Cenomanian
Chartense amber
France
An oonopid
Orchestina rabagensis [ 11]
Species
Valid
Saupe et al.
Early Albian
Spain
An oonopid, a species of Orchestina .
Palaeogrosphus jacquesi [ 12]
Sp. nov
Valid
Lourenço & Henderickx
Holocene
Malagasy copal
Madagascar
A buthid scorpion .
Palaeoleptoneta [ 2]
Gen. et sp. nov
Valid
Wunderlich
Cretaceous
Burmese amber
Myanmar
Originally described as a member of Leptonetidae , subsequently considered to be a member of Araneomorphae of uncertain affinities by Magalhaes et al. .[ 13] The type species is P. calcar .
Palaeolychas weitschati [ 14]
Sp. nov.
Valid
Lourenço
Eocene
Russia (Kaliningrad Oblast )
A buthid scorpion , a species of Palaeolychas .
Palaeotrilineatus [ 15]
Gen. et sp. nov.
Valid
Lourenço
Early Cretaceous
Burmese amber
Myanmar
A scorpion . The type species is P. ellenbergeri .
?Pholcochyrocer baculum [ 2]
Sp. nov
Valid
Wunderlich
Cretaceous
Burmese amber
Myanmar
A member of Pholcochyroceridae , possibly a species of Pholcochyrocer .
Pholcochyrocer pecten [ 2]
Sp. nov
Valid
Wunderlich
Cretaceous
Burmese amber
Myanmar
A member of Pholcochyroceridae , a species of Pholcochyrocer .
Piankhi [ 16]
Gen. et sp. nov.
Valid
Dunlop, Bartel & Mitov
Eocene
Baltic amber
Europe
A harvestman found in Baltic amber . The type species is P. steineri .
Piankhi steineri
?Poliochera cretacea [ 17]
Sp. nov
Valid
Wunderlich
Cretaceous
Burmese amber
Myanmar
A member of Ricinulei , possibly a species of Poliochera .
Praoppiella [ 18]
Gen. et sp. nov.
Valid
Miko & Mourek in Miko et al.
Early Pleistocene
Slovenia
A mite . The type species is P. oanae .
Pseudogarypus synchrotron [ 19]
Sp. nov
Valid
Henderickx in Henderickx, Tafforeau & Soriano
Late Eocene
Baltic Amber
Europe
A pseudoscorpion
Pseudogarypus synchrotron
?Psiloderces filiformis [ 2]
Sp. nov
Valid
Wunderlich
Cretaceous
Burmese amber
Myanmar
First described as possibly species of Psiloderces ; transferred to the genus Eopsiloderces .[ 8]
Rhinoppioides [ 18]
Gen. et sp. nov.
Valid
Miko in Miko et al.
Early Pleistocene
Slovenia
A mite . The type species is R. quadrituberculatus .
Saetosoma [ 2]
Gen. et sp. nov
Valid
Wunderlich
Cretaceous
Burmese amber
Myanmar
A member of Tetrablemmidae . The type species is S. filiembolus .
Samlandicmeta [ 3]
Gen. et sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
A long-jawed orb weaver . The type species is S. mutila .
?Scytodes hani [ 2]
Sp. nov
Valid
Wunderlich
Cretaceous
Lebanese amber
Jordan
A spider of uncertain affinities. Originally described as a spitting spider , possibly a species of Scytodes ; Magalhaes et al. subsequently argued that it cannot be reliably assigned to Scytodoidea .[ 13]
Succiniropsis runcinata [ 3]
Sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
A zoropsid .
?Succinomus gibbosus [ 3]
Sp. nov
Valid
Wunderlich
Eocene
Baltic amber
Europe
A succinomid , possibly a species of Succinomus .
Trhypochthonius lopezvallei [ 20]
Sp. nov.
Valid
Arillo, Subías & Shtanchaeva
Early Cretaceous (Albian)
Spain
A trhypochthoniid mite .
Triasacarus [ 1]
Gen. et sp. nov.
Valid
Lindquist & Grimaldi in Schmidt et al.
Carnian
Italy
A mite . The type species is T. fedelei .
Crustaceans
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Acanthocythereis inouei [ 21]
Sp. nov
Valid
Yamaguchi, Mashiba & Kamiya
Miocene
Osaki Formation
Japan
An ostracod , a species of Acanthocythereis .
Acanthophoenicides [ 22]
Gen. et sp. nov
Valid
Audo & Charbonnier
Late Cretaceous (Cenomanian )
Hadjoula Lagerstätte
Lebanon
A slipper lobster belonging to the subfamily Arctidinae. The type species is A. peterpani .
Acratia candyae [ 23]
Sp. nov
Valid
Forel
Late Permian (Changhsingian )
Wujiaping Formation
China
An acratiid ostracod , a species of Acratia .
Agnocarcinus [ 24]
Gen. et sp. nov
Valid
Beschin et al.
Eocene (Lutetian )
Italy
A crab belonging to the family Pilumnidae . The type species is A. zannatoi .
Alberticarcinus [ 24]
Gen. et sp. nov
Valid
Beschin et al.
Eocene (Lutetian )
Italy
A crab belonging to the family Atelecyclidae . The type species is A. eocaenus .
Ammopylocheles robertboreki [ 25]
Sp. nov
Valid
Fraaije et al.
Middle Oxfordian
Poland
A pylochelid , a species of Ammopylocheles .
Amphicytherura grandicribra [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A schizocytherid ostracod , a species of Amphicytherura .
Amphicytherura occulta [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
An ostracod . Originally described as a species of Amphicytherura ; Antonietto et al. (2013) transferred this species to the genus Dinglecythere .[ 27]
Amphissites gifuensis [ 28]
Sp. nov
Valid
Tanaka & Yuan in Tanaka et al.
Early Permian
Japan
An ostracod , a species of Amphissites .
Antonioranina [ 29]
Gen. et comb. nov
Valid
Van Bakel et al.
Eocene
Italy
Spain
United States
A raninid crab . The type species is "Cyrtorhina " globosa Beschin, Busulini, De Angeli & Tessier (1988); genus also contains A. fusseli (Blow & Manning, 1996) and A. ripacurtae (Artal & Castillo, 2005). Van Bakel et al. (2012) tentatively assigned "Cyrtorhina" oblonga Beschin, Busulini, De Angeli & Tessier (1988) to the genus Antonioranina as well;[ 29] however, Karasawa et al. (2014) made it the type species of a separate genus Claudioranina .[ 30]
Archaeoniscus coreaensis [ 31]
Sp. nov
Valid
Park et al.
Early Cretaceous
Jinju Formation
South Korea
An isopod crustacean, a species of Archaeoniscus .
Asciocythere cabbagehillensis [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A cytherideid ostracod , a species of Asciocythere .
Asymmetricythere semoupathei [ 32]
Sp. nov
Valid
Sarr
Ypresian to Lutetian
Mauritania
Senegal
A trachyleberidid ostracod , a species of Asymmetricythere .
Aurikirkbya miyakei [ 28]
Sp. nov
Valid
Tanaka & Ono in Tanaka et al.
Early Permian
Japan
An ostracod , a species of Aurikirkbya .
Aysegulina chapeltonensis [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A hemicytherid ostracod , a species of Aysegulina .
Aysegulina riominhoensis [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A hemicytherid ostracod , a species of Aysegulina .
Aysegulina sagitta [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A hemicytherid ostracod , a species of Aysegulina .
Aysegulina ventrocurva [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A hemicytherid ostracod , a species of Aysegulina .
Bairdia adelineae [ 23]
Sp. nov
Valid
Forel
Early Triassic (Griesbachian )
Daye Formation
China
A bairdiid ostracod , a species of Bairdia .
Bairdia fujisan [ 28]
Sp. nov
Valid
Tanaka & Maeda in Tanaka et al.
Early Permian
Japan
An ostracod , a species of Bairdia .
Bairdia? huberti [ 23]
Sp. nov
Valid
Forel
Late Permian (Changhsingian )
Wujiaping Formation
China
A bairdiid ostracod , a possible species of Bairdia .
Bairdia ikeyai [ 28]
Sp. nov
Valid
Tanaka in Tanaka et al.
Early Permian
Japan
An ostracod , a species of Bairdia .
Bairdia jeromei [ 23]
Sp. nov
Valid
Forel
Late Permian (Changhsingian ) and Early Triassic (Griesbachian )
Daye Formation
Wujiaping Formation
China
A bairdiid ostracod , a species of Bairdia .
Bairdia minoensis [ 28]
Sp. nov
Valid
Tanaka & Ono in Tanaka et al.
Early Permian
Japan
An ostracod , a species of Bairdia .
Bairdia spindlica [ 28]
Sp. nov
Valid
Tanaka & Yuan in Tanaka et al.
Early Permian
Japan
An ostracod , a species of Bairdia .
Balowella crassicostata [ 33]
Sp. nov
Valid
Luppold
Middle Jurassic
Germany
An ostracod , a species of Balowella .
Bathypluma pliocenica [ 34]
Sp. nov
Valid
Garassino, Pasini & Marini
Early Pliocene
Italy
A retroplumid crab , a species of Bathypluma .
Bournelyreidus [ 29]
Gen. et comb. et sp. nov
Valid
Van Bakel et al.
Late Cretaceous to Paleocene
Denmark
France
Germany
Greenland
United States
A lyreidid crab . The type species is "Hemioon " eysunesensis Collins & Wienberg Rasmussen (1992); according to the authors of its description genus also contains "Raninella " carlilensis Feldmann & Maxey (1980), "Palaeocorystes " laevis Schlüter in von der Marck & Schlüter (1868), "Raninella" manningi Bishop & Williams (2000), "Raninella" oaheensis Bishop (1978) and "Raninella" tridens Roberts (1962), as well as the new species Bournelyreidus teodorii .[ 29] On the other hand, Karasawa et al. (2014) transferred B. teodorii to the genus Lyreidus , "R." tridens to the genus Macroacaena and "R." manningi to the genus Heus .[ 30]
Brachycythere jamaicaensis [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A trachyleberidid ostracod , a species of Brachycythere .
Bretonia [ 35]
gen et comb nov
Valid
Schweitzer et al.
Cenomanian
Sables du Perche Formation
France
crab , New genus for "Xanthosia " danielae (Collins and Breton, 2009).
Buntonia bodergati [ 32]
Sp. nov
Valid
Sarr
Thanetian to Ypresian
Mali
Senegal
A trachyleberidid ostracod , a species of Buntonia .
Buntonia nana [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A trachyleberidid ostracod , a species of Buntonia .
Buntonia vulgaris [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A trachyleberidid ostracod , a species of Buntonia .
Bythocypris antoniettae [ 36]
Sp. nov
Valid
Sciuto
Early Pleistocene
Italy
An ostracod , a species of Bythocypris .
Bythocythere solisdeus [ 37]
Sp. nov
Valid
Sciuto
Early Pleistocene
Italy
A bythocytherid ostracod , a species of Bythocythere .
Calliaxina altavillensis [ 38]
Sp. nov
Valid
Beschin & De Angeli
Eocene
Italy
A member of Callianassidae .
Carcinoplax parsensis [ 39]
Sp. nov
Valid
Vega et al.
Miocene (Burdigalian or Langhian )
Mishan Formation
Iran
A goneplacid decapod crustacean, a species of Carcinoplax .
Carpopenaeus garassinoi [ 40]
Sp. nov
Valid
Charbonnier in Petit & Charbonnier
Late Cretaceous
Sahel Alma Lagerstätte
Lebanon
Cenocorystes bretoni [ 29]
Sp. nov
Valid
Van Bakel et al.
Late Cretaceous (late Cenomanian)
France
A palaeocorystid crab , a species of Cenocorystes .
Cenomanocarcinus cantabricus [ 29]
Sp. nov
Valid
Van Bakel et al.
Early Cretaceous (Albian)
Spain
A cenomanocarcinid crab , a species of Cenomanocarcinus .
Coleia martinlutheri [ 41]
sp nov
Valid
Schweigert & Ernst
Sinemurian
Germany
A coleiid polychelid , a species of Coleia .
Cretacolana [ 42]
Gen. et comb. nov
Valid
Schweitzer & Feldmann
Cenomanian
France
A porcelain crab , a new genus for "Porcellana" antiqua A. Milne-Edwards, 1862.
Cretatrizocheles [ 43]
gen et sp nov
Valid
Fraaije, Klompmaker, & Artal
Albian or Cenomanian
Spain
A hermit crab . The type species is Cretatrizocheles olazagutiensis .
Cromimaia [ 24]
Gen. et comb. nov
Valid
Beschin et al.
Eocene
Italy
A crab belonging to the family Majidae . The type species is "Micromaia " meneguzzoi Beschin, Busulini, De Angeli & Tessier (1985); genus also includes "Micromaia" elegans Beschin, Busulini, De Angeli & Tessier (1985).
Cushmanidea tessieri [ 32]
Sp. nov
Valid
Sarr
Ypresian to Lutetian
Senegal
A cushmanideid ostracod , a species of Buntonia .
Cytherelloidea guineacornensis [ 26]
Sp. nov
Valid
Puckett in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Guinea Corn Formation
Jamaica
A cytherellid ostracod , a species of Cytherelloidea .
Cytheropteron eleonorae [ 37]
Sp. nov
Valid
Sciuto
Early Pleistocene
Italy
A cytherurid ostracod , a species of Cytheropteron .
Cytheropteron italoi [ 36]
Sp. nov
Valid
Sciuto
Early Pleistocene
Italy
An ostracod , a species of Cytheropteron .
Cytheropteron rossanae [ 36]
Sp. nov
Valid
Sciuto
Early Pleistocene
Italy
An ostracod , a species of Cytheropteron .
Daira coccoi [ 24]
Sp. nov
Valid
Beschin et al.
Eocene (Lutetian )
Italy
A crab belonging to the family Dairidae ; a species of Daira .
Dinochelus steeplensis [ 44]
Sp. nov
Valid
Tshudy & Saward
Eocene (Ypresian )
London Clay
United Kingdom
A lobster , a species of Dinochelus .
Distefania renefraaijei [ 45]
Sp. nov
Valid
Klompmaker, Feldmann & Schweitzer
Cretaceous (Albian or Cenomanian)
Spain
A goniodromitid crab , a species of Distefania .
Dolocythere tuberculata [ 33]
Sp. nov
Valid
Luppold
Middle Jurassic
Germany
An ostracod . Originally described as a species of Dolocythere ; Lord, Cabral & Danielopol (2020) transferred this species to the genus Minyocythere .[ 46]
Ebullitiocaris elatus [ 47]
Sp. nov
Valid
Womack et al.
Carboniferous
United Kingdom
A member of Cladocera , a species of Ebullitiocaris .
Eocytheropteron hazeli [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A cytherurid ostracod , a species of Eocytheropteron .
Eocytheropteron jamaicaensis [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A cytherurid ostracod , a species of Eocytheropteron .
Eouroptychus [ 48]
Gen. et sp. nov
Valid
De Angeli & Ceccon
Eocene (Ypresian )
Italy
A member of Chirostylidae . The type species is E. montemagrensis .
Eucytherura mitchelli [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A cytherurid ostracod , a species of Eucytherura .
Eomunidopsis aldoirarensis [ 49]
Sp. nov
Valid
Klompmaker et al.
Cretaceous (Albian or Cenomanian)
Spain
A galatheoid , a species of Eomunidopsis .
Eopaguropsis nidiaquilae [ 50]
Sp. nov
Valid
Fraaije et al.
Oxfordian
Poland
A diogenid , a species of Eopaguropsis .
Eosymethis [ 29]
Gen. et sp. nov
Valid
Van Bakel et al.
Eocene (Ypresian )
Roda Formation
Spain
A raninid crab . The type species is Eosymethis aragonensis .
Eohexapus orthogonius [ 24]
Sp. nov
Valid
Beschin et al.
Eocene (Lutetian )
Italy
A crab belonging to the family Hexapodidae .
Eucorystes iserbyti [ 29]
Sp. nov
Valid
Van Bakel et al.
Early Cretaceous (Albian)
France
A palaeocorystid crab , a species of Eucorystes .
Eucorystes navarrensis [ 29]
Sp. nov
Valid
Van Bakel et al.
Early Cretaceous (Albian)
Spain
A palaeocorystid crab , a species of Eucorystes .
Faksecarcinus [ 35]
gen et comb nov
Valid
Schweitzer et al.
Albian or Cenomanian to Danian
Denmark
Spain
A crab, a new genus for "Xanthosia" gracilis .
Ferroranina [ 29]
Gen. et comb. et sp. nov
Valid
Van Bakel et al.
Late Cretaceous (late Cenomanian to late Campanian)
Czech Republic [ 51]
India
Madagascar
Mexico
United States
A palaeocorystid crab . The type species is "Notopocorystes " dichrous Stenzel (1945); genus also contains "Notopocorystes" australis Secretan (1964) and "Notopocorystes" fritschi Glaessner (1929),[ 51] as well as the new species Ferroranina tamilnadu .
Floricythereis [ 26]
Gen. et sp. et comb. nov
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
United States
A trachyleberidid ostracod . Genus includes new species Floricythereis exquisita , as well as F. lixula (Crane, 1965) (the type species was not designated).
Galathea mainensis [ 52]
Sp. nov
Valid
Ceccon & De Angeli
Middle Eocene
Italy
A species of Galathea .
Gebiacantha tuscia [ 53]
Sp. nov
Valid
Garassino, Pasini, De Angeli & Charbonnier in Garassino et al.
Early Pliocene
Italy
An upogebiid decapod, a species of Gebiacantha .
Geffenina mariebeatriceae [ 54]
Sp. nov
Valid
Chitnarin in Chitnarin et al.
Late Early Permian
Tak Fa Formation
Thailand
A knoxitid palaeocopidan ostracod , a species of Geffenina .
Geffenina mariebeatriceae [ 54]
Sp. nov
Valid
Chitnarin in Chitnarin et al.
Early Permian
Pha Nok Khao Formation
Thailand
A knoxitid palaeocopidan ostracod , a species of Geffenina .
Glyphea wiffenae [ 55]
Species
Valid
Feldmann and Stilwell
Late Cretaceous
Maungataniwa Sandstone
New Zealand
A glypheid lobster, a species of Glyphea .
Glyptopleura mugiensis [ 28]
Sp. nov
Valid
Tanaka & Yuan in Tanaka et al.
Early Permian
Japan
An ostracod , a species of Glyptopleura .
Goniodromites kubai [ 56]
sp nov
Valid
Starzyk, Krzemiska, & Krzemiski
Oxfordian
Poland
A crab , a species of Goniodromites .
Gonioplacoides [ 57]
Gen. et sp. nov
Valid
Quayle & Collins
Eocene
Headon Hill Formation
United Kingdom
A goneplacid . The type species is Gonioplacoides minuta .
Graptocarcinus urbasaensis [ 58]
Sp. nov
Valid
Van Bakel et al.
Late Cretaceous
Spain
A dynomenid crab , a species of Graptocarcinus .
Grinioneis pachycosta [ 59]
Sp. nov
Valid
Guernet et al.
Eocene (Lutetian )
Paris Basin
France
A trachyleberidid ostracod , a species of Grinioneis .
Grolamaia [ 24]
Gen. et sp. nov
Valid
Beschin et al.
Eocene (Lutetian )
Italy
A crab belonging to the subfamily Pisinae . The type species is G. vicariottoi .
Hadrocarcinus [ 60]
gen et comb et sp nov
Valid
Schweitzer, Feldmann & Lamanna
Late Cretaceous
Antarctica
A necrocarcinid crab, a new genus for "Necrocarcinus " wrighti . The genus also contains "Necrocarcinus" carinatus and a new species Hadrocarcinus tectilacus .
Harbinia alta [ 61]
Sp. nov
Valid
Antonietto et al.
Aptian or early Albian
Santana Formation
Brazil
An ostracod crustacean, a species of Harbinia .
Headonipus [ 57]
Gen. et sp. nov
Valid
Quayle & Collins
Late Eocene
Headon Beds
United Kingdom
A hexapodid . The type species is Headonipus tuberculatus .
Hemiparacytheridea exquisita [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A paracytherideid ostracod , a species of Hemiparacytheridea .
Hispanigalathea [ 49]
Gen. et 2 sp. nov
Valid
Klompmaker et al.
Cretaceous (Albian or Cenomanian)
Spain
A galatheoid . Genus contains two species: Hispanigalathea pseudolaevis and H. tuberosa .
Huanghestheria linxiensis [ 62]
Sp. nov
Valid
Niu in Zhang et al.
Late Permian
Linxi Formation
China
A vertexiine lioestheriid spinicaudatan clam shrimp , a species of Huanghestheria .
Joeranina [ 29]
Gen. et comb. et sp. nov
Valid
Van Bakel et al.
Cretaceous (Albian to early Campanian)
Canada
France
Greenland
Japan
Madagascar
Spain
Switzerland
Syria
United Kingdom
United States
A palaeocorystid crab . The type species is "Corystes " broderipii Mantell (1844). According to the authors of its description genus also contains "Palaeocorystes " harveyi Woodward (1896), "Eucorystes " japonicus Jimbô (1894), "Notopocorystes " paututensis Collins & Wienberg Rasmussen (1992), "Eucorystes" platys Schweitzer & Feldmann (2002) and "Notopocorystes" syriacus Withers (1928), as well as the new species Joeranina gaspari .[ 29] On the other hand, Karasawa et al. (2014) transferred "P." harveyi , E. platys and "N." paututensis to the genus Eucorystes ; the authors also transferred the species "Notopocorystes" xizangensis Wang (1981) to the genus Joeranina .[ 30]
Juracrista [ 63]
Gen. et 2 sp. nov
Valid
Robins, Feldmann & Schweitzer
Late Jurassic (Tithonian)
Ernstbrunn Limestone
Austria
A munidid . The type species is Juracrista perculta ; genus also contains Juracrista costaspinosa .
Jurapylocheles iwonae [ 25]
Sp. nov
Valid
Fraaije et al.
Middle Oxfordian
Poland
A pylochelid , a species of Jurapylocheles .
Kinkelinella circumcostata [ 33]
Sp. nov
Valid
Luppold
Middle Jurassic
Germany
An ostracod , a species of Kinkelinella .
Koryncheiros [ 64]
Gen. et sp. nov
Valid
Feldmann et al.
Middle or late Anisian
Guanling Formation
China
A decapod crustacean; a clytiopsid erymoid . The type species is Koryncheiros luopingensis .
Kroemmelbeincypris [ 65]
Gen. et comb. nov
Valid
Poropat & Colin
Aptian
An ostracod crustacean, a new genus for the species "Hourcqia " symmetrica Krömmelbein & Weber and "Hourcqia" angulata Krömmelbein & Weber.
Langdaia meesooki [ 54]
Sp. nov
Valid
Chitnarin in Chitnarin et al.
Late Early to Middle Permian
Tak Fa Formation
Thailand
A knoxitid palaeocopidan ostracod , a species of Langdaia .
Leiocythereis [ 26]
Gen. et sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A trachyleberidid ostracod . The type species is Leiocythereis polita .
Lepas sattmanni [ 66]
Sp. nov
Valid
Harzhauser & Schlögl
Early Miocene
Ebelsberg Formation
Austria
A lepadid barnacle , a species of Lepas .
Lessinicarcinus [ 67]
Gen. et comb. nov
Valid
De Angeli
Eocene
Croatia [ 68]
Italy
A crab belonging to the family Pilumnidae . The type species is "Titanocarcinus " euglyphos Bittner (1875).
Lessinipagurus [ 24]
Gen. et sp. nov
Valid
Beschin et al.
Eocene (Lutetian )
Italy
A hermit crab . Originally assigned to the family Paguridae ,[ 24] subsequently transferred to the separate family Xylopaguridae by Gašparič et al. (2016).[ 69] The type species is L. granulatus .
Lienenklausicythere [ 70]
Gen. et comb. nov
Valid
Uffenorde, Monostori & Tóth
Eocene to Oligocene
Europe
An ostracod , a new genus for "Hermanites " acuticosta and "Hermanites " camela ; genus also contains a new, yet-unnamed species.
Ljubimovella debilicostatus [ 33]
Sp. nov
Valid
Luppold
Middle Jurassic
Germany
An ostracod , a species of Ljubimovella .
Lobonotus beschini [ 71]
Sp. nov
Valid
De Angeli & Checchi
Eocene (Lutetian )
Italy
A crab belonging to the family Tumidocarcinidae .
Loxoconcha pindarsensis [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A loxoconchid ostracod , a species of Loxoconcha .
Marylyreidus [ 29]
Gen. et comb. nov
Valid
Van Bakel et al.
Cretaceous (late Albian–early Cenomanian)
Pawpaw Formation
United States
A lyreidid crab . The type species is "Notopocorystes " punctatus Rathbun (1935).
Megokkos patagoniensis [ 72]
Sp. nov
Valid
Schweitzer et al.
Eocene
Rio Turbio Formation
Argentina
A crab , a species of Megokkos .
Mesoparapylocheles [ 43]
gen et sp nov
Valid
Fraaije, Klompmaker, & Artal
Albian or Cenomanian
Spain
A hermit crab. The type species is Mesoparapylocheles michaeljacksoni .
Microcheilinella subovata [ 28]
Sp. nov
Valid
Tanaka & Maeda in Tanaka et al.
Early Permian
Japan
An ostracod , a species of Microcheilinella .
Microcheilinella tsubogawensis [ 28]
Sp. nov
Valid
Tanaka & Ichida in Tanaka et al.
Early Permian
Japan
An ostracod , a species of Microcheilinella .
Microcoelonella takfaensis [ 54]
Sp. nov
Valid
Chitnarin in Chitnarin et al.
Middle Permian
Tak Fa Formation
Thailand
A coelonellid palaeocopidan ostracod , a species of Microcoelonella .
Microcoelonella takliensis [ 54]
Sp. nov
Valid
Chitnarin in Chitnarin et al.
Middle Permian
Tak Fa Formation
Thailand
A coelonellid palaeocopidan ostracod , a species of Microcoelonella .
Munida deangelii [ 73]
Sp. nov
Valid
Robins, Feldmann & Schweitzer
Late Oligocene to early Miocene
Río Foyel Formation
Argentina
A munidid , a species of Munida .
Myra gurii [ 39]
Sp. nov
Valid
Vega et al.
Miocene (Burdigalian or Langhian )
Mishan Formation
Iran
A leucosiid decapod crustacean, a species of Myra .
Navarradromites [ 45]
Gen. et sp. nov
Valid
Klompmaker, Feldmann & Schweitzer
Cretaceous (Albian or Cenomanian)
Spain
A goniodromitid crab . The type species is Navarradromites pedroartali .
Navarrahomola [ 74]
Gen. et sp. nov.
Valid
Artal et al.
Cretaceous (Albian or Cenomanian)
Spain
A homolid crab . The type species is Navarrahomola hispanica .
Neocallichirus bacatus [ 75]
Sp. nov
Valid
Busulini, Beschin & Tessier
Eocene (Priabonian )
Italy
A member of Callianassidae .
Neocytherideis labyrinthoidea [ 59]
Sp. nov
Valid
Guernet et al.
Eocene (Lutetian )
Paris Basin
France
A neocytherideid ostracod , a species of Neocytherideis .
Neocytheromorpha priscipacifica [ 21]
Sp. nov
Valid
Yamaguchi, Mashiba & Kamiya
Miocene
Osaki Formation
Japan
An ostracod , a species of Neocytheromorpha .
Neurocythere jakovlevae [ 76]
Sp. nov
Valid
Kolpenskaya in Tesakova et al.
Late Jurassic (Early Kimmeridgian)
Russia
An ostracod , a species of Neurocythere .
Nitotacarcinus antipodes [ 72]
Sp. nov
Valid
Schweitzer et al.
Eocene
Rio Turbio Formation
Argentina
A crab , a species of Nitotacarcinus .
Nodosculda [ 77]
Gen. et sp. nov
Valid
Franţescu
Early Cretaceous (late Albian)
United States
A mantis shrimp . The type species is Nodosculda fisherorum .
Noetlingocarcinus messinai [ 24]
Sp. nov
Valid
Beschin et al.
Eocene (Lutetian )
Italy
A crab belonging to the superfamily Dromioidea and the family Basinotopidae , a species of Noetlingocarcinus .
Nogarhomola [ 78]
Gen. et sp. nov
Valid
De Angeli & Alberti
Eocene (Lutetian )
Italy
A homolid crab . The type species is Nogarhomola aurorae .
Notopocorystes kerri [ 79]
Sp. nov
Valid
Luque et al.
Early Cretaceous (late Aptian)
Paja Formation
Colombia
A palaeocorystid raninoid crab . Initially described as a species of Notopocorystes ,[ 79] but subsequently reassigned to the genus Joeranina .[ 80]
Nykteripteryx [ 49]
Gen. et sp. nov
Valid
Klompmaker et al.
Cretaceous (Albian or Cenomanian)
Spain
A galatheoid . The type species is Nykteripteryx rostrata .
Ochtholambrus gambillarai [ 75]
Sp. nov
Valid
Busulini, Beschin & Tessier
Eocene (Priabonian )
Italy
A crab belonging to the family Parthenopidae .
Odontodactylus italicus [ 24]
Sp. nov
Valid
Beschin et al.
Eocene (Lutetian )
Italy
A mantis shrimp belonging to the family Odontodactylidae; a species of Odontodactylus .
Ommaciria [ 24]
Gen. et comb. nov
Valid
Beschin et al.
Eocene
Italy
A crab belonging to the family Majidae . The type species is "Micromaia " mainensis Beschin, Busulini, De Angeli & Tessier (1985).
Orthobairdia jeanlouisi [ 23]
Sp. nov
Valid
Forel
Early Triassic (Griesbachian )
Daye Formation
China
A bairdiid ostracod , a species of Orthobairdia .
Ovocytheridea rotunda [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A cytherideid ostracod , a species of Ovocytheridea .
Paguristes gagnaisoni [ 81]
Sp. nov
Valid
Gagnaison
Miocene
France
A species of Paguristes .
Pagurus valdagnensis [ 24]
Sp. nov
Valid
Beschin et al.
Eocene (Lutetian )
Italy
A hermit crab belonging to the family Paguridae , a species of Pagurus .
Palaega picena [ 82]
Sp. nov
Valid
Pasini & Garassino
Miocene
Italy
A cirolanid isopod , a species of Palaega .
Palaega pisana [ 83]
Sp. nov
Junior synonym
Pasini & Garassino
Pliocene
Italy
A cirolanid isopod . Originally described as a species of Palaega ; considered to be a junior synonym of Bathynomus sismondai (Ristori, 1891) by Hyžný, Pasini & Garassino (2019).[ 84]
Palaega steatopigia [ 85]
Sp. nov
Nomen dubium
Pasini & Garassino
Pliocene
Italy
A cirolanid isopod . Originally described as a species of Palaega ; Hyžný, Pasini & Garassino (2019) considered this species to be a nomen dubium and reassigned it to the genus Bathynomus .[ 84]
Palaeohomarus pacificus [ 86]
Sp. nov
Valid
Aguirre-Urreta, Lazo & Rawson
Late Hauterivian or early Barremian
Agrio Formation
Argentina
A decapod crustacean, a species of Palaeohomarus .
Paracypris keiji [ 59]
Sp. nov
Valid
Guernet et al.
Eocene (Lutetian )
Paris Basin
France
A candonid ostracod , a species of Paracypris .
Paranecrocarcinus balla [ 29]
Sp. nov
Valid
Van Bakel et al.
Late Cretaceous (Cenomanian)
France
A necrocarcinid crab , a species of Paranecrocarcinus .
Periosocypris [ 87]
Gen. et sp. nov
Valid
Whatley, Khosla & Rathore
Late Cretaceous (Maastrichtian)
Lameta Formation
India
A cypridid ostracod . The type species is Periosocypris megistus .
Petrochirus savii [ 24]
Sp. nov
Valid
Beschin et al.
Eocene (Lutetian )
Italy
A hermit crab belonging to the family Diogenidae , a species of Petrochirus .
Phrixocythere [ 26]
Gen. et sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A hemicytherid ostracod . The type species is Phrixocythere unionensis .
Planocarcinus [ 79]
Gen. et comb. nov
Valid
Luque et al.
Early Cretaceous (late Aptian)
Paja Formation
Colombia
A necrocarcinid raninoid crab , a new genus for "Dakoticancer " olssoni Rathbun (1937).
Podocallichirus laepaensis [ 88]
Sp. nov
Valid
Hyžný & Muñiz
Late Miocene
Spain
A callianassid ghost shrimp . Originally described as a species of Podocallichirus ; subsequently transferred to the genus Balsscallichirus by Hyžný (2016).[ 89]
Protobuntonia warorei [ 32]
Sp. nov
Valid
Sarr
Paleocene to Ypresian
Senegal
Togo
A trachyleberidid ostracod , a species of Buntonia .
Pterygocythereis babinoti [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A trachyleberidid ostracod , a species of Pterygocythereis .
Pylochelitergites [ 90]
Gen. et 2 sp. nov
Valid
Fraaije et al.
Oxfordian to Kimmeridgian
Germany
Poland
A pylochelid hermit crab . Genus contains two species: P. westerbergensis (the type species) and P. gelasinus .
Quasilaeviranina minuscula [ 24]
Sp. nov
Valid
Beschin et al.
Eocene (Lutetian )
Italy
A crab belonging to the family Raninidae , a species of Quasilaeviranina .
Raninoides rioturbiensis [ 72]
Sp. nov
Valid
Schweitzer et al.
Eocene
Rio Turbio Formation
Argentina
A crab , a species of Raninoides .
Rhinolambrus elongatus [ 24]
Sp. nov
Valid
Beschin et al.
Eocene (Lutetian )
Italy
A crab belonging to the family Parthenopidae ; a species of Rhinolambrus .
Ristoria [ 53]
Gen. et comb. nov
Valid
Garassino, Pasini, De Angeli & Charbonnier in Garassino et al.
Early Pliocene
Italy
A leucosiid crab , a new genus for "Ilia " pliocaenica Ristori, 1891.
Samarella sonei [ 54]
Sp. nov
Valid
Chitnarin in Chitnarin et al.
Middle Permian
Tak Fa Formation
Thailand
A paraparchitid palaeocopidan ostracod , a species of Samarella .
Samarella viscusforma [ 54]
Sp. nov
Valid
Chitnarin in Chitnarin et al.
Early to Middle Permian
Pha Nok Khao Formation
Tak Fa Formation
Thailand
A paraparchitid palaeocopidan ostracod , a species of Samarella .
Sargentina chantarameei [ 54]
Sp. nov
Valid
Chitnarin in Chitnarin et al.
Early to Middle Permian
Pha Nok Khao Formation
Tak Fa Formation
Thailand
A knoxitid palaeocopidan ostracod , a species of Sargentina .
Scalpellum paratethyianum [ 66]
Sp. nov
Valid
Harzhauser & Schlögl
Early Miocene
Lakšárska Nová Ves Formation
Slovakia
A scalpellid barnacle , a species of Scalpellum .
Schizoptocythere nana [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A trachyleberidid ostracod , a species of Schizoptocythere .
Schizoptocythere pinna [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A trachyleberidid ostracod , a species of Schizoptocythere .
Schuleridea exigua [ 26]
Sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
A cytherideid ostracod , a species of Schuleridea .
Semicytherura carioui [ 32]
Sp. nov
Valid
Sarr
Ypresian to Lutetian
Senegal
A cytherurid ostracod , a species of Semicytherura .
Sineruga [ 91]
gen et sp nov
Valid
Silurian (uppermost Ludlow to lower Pridoli)
Le Val Formation
France
myodocope ostracod . type species is Sineruga insolita .
Southcavea (Saxosouthcavea) [ 33]
Subgen. et 3 sp. nov
Valid
Luppold
Middle Jurassic
Germany
An ostracod , a subgenus of Southcavea containing three new species: Southcavea (Saxosouthcavea) extendata , Southcavea (Saxosouthcavea) bartenbrandi and Southcavea (Saxosouthcavea) malzi (with two subspecies: S. (S.) malzi malzi and S. (S.) malzi reticulata ).
Spinicytheridea [ 26]
Gen. et sp. nov
Valid
Puckett & Colin in Puckett, Colin & Mitchell
Late Cretaceous (Maastrichtian )
Jamaica
Mexico
A cytherideid ostracod . The type species is Spinicytheridea compta from Jamaica; genus also includes a second, unnamed species from the Maastrichtian Ocozocoautla Formation of Chiapas, Mexico.
Spinirostrimaia [ 24]
Gen. et comb. nov
Valid
Beschin et al.
Eocene
Italy
Spain
A crab belonging to the family Majidae . The type species is "Micromaia " margaritata Fabiani (1910).
Stagmacaris adielklompmakeri [ 90]
Sp. nov
Valid
Fraaije et al.
Early late Kimmeridgian
Germany
A pylochelid hermit crab , a species of Stagmacaris .
Stagmacaris biarcus [ 90]
Sp. nov
Valid
Fraaije et al.
Middle Oxfordian
Poland
A pylochelid hermit crab , a species of Stagmacaris .
Steorrosia [ 35]
gen et comb nov
Valid
Schweitzer et al.
Late Albian to Cenomanian
United States
Crab, New genus for "Xanthosia" aspera . Other species S. pawpawensis and S. reidi .
Styracocarcinus [ 42]
Gen. et comb. nov
Valid
Schweitzer & Feldmann
Late Cretaceous (Senonian)
Morocco
A tumidocarcinid decapod , a new genus for "Titanocarcinus " meridionalis Secretan, 1961.
Superpendicythere [ 33]
Gen. et sp. nov
Valid
Luppold
Middle Jurassic
Germany
An ostracod . The type species is S. binodosa .
Symethoides [ 29]
Gen. et sp. nov
Valid
Van Bakel et al.
Paleocene (early Danian )
Tinton Formation
United States
A raninid crab . The type species is Symethoides monmouthorum .
Szaboa deppermanni [ 92]
Sp. nov
Valid
Fraaije et al.
Oligocene
Ratzeburg Formation
Germany
A matutid crab, a species of Szaboa .
Thalamita italica [ 53]
Sp. nov
Valid
Garassino, Pasini, De Angeli & Charbonnier in Garassino et al.
Early Pliocene
Italy
A portunid crab , a species of Thalamita .
Tridactylastacus [ 64]
Gen. et sp. nov
Valid
Feldmann et al.
Middle or late Anisian
Guanling Formation
China
A decapod crustacean; a glypheoid belonging to the family Litogastridae .[ 93] The type species is Tridactylastacus sinensis .
Turbiocheir [ 72]
Gen. et sp. nov
Disputed
Schweitzer et al.
Eocene
Rio Turbio Formation
Argentina
Chile
Venezuela [ 94]
A thalassinidean . The type species is Turbiocheir minutospinata . Hyžný et al. (2020) considered the genus Turbiocheir to be a junior synonym of the genus Anacalliax , creating a new combination Anacalliax minutospinata .[ 95]
Vecticallichirus [ 57]
Gen. et sp. nov
Valid
Quayle & Collins
Late Eocene and Oligocene[ 96]
Headon Beds
United Kingdom
A callichirid [ 96] ghost shrimp. The type species is Vecticallichirus abditus . Hyžný (2020) considered V. abditus to be a junior synonym of "Callianassa " batei Woodward (1869), resulting in a new combination Vecticallichirus batei .[ 96]
Vegaranina [ 29]
Gen. et comb. nov
Valid
Van Bakel et al.
Late Cretaceous (Campanian–Maastrichtian)
Cuba
France
Mexico
United States ( Puerto Rico )
A raninid crab . The type species is "Lophoranina " precocious Feldmann, Vega, Tucker, García-Barrera & Avendaño (1996).
Vernoniella tenuipora [ 33]
Sp. nov
Valid
Luppold
Middle Jurassic
Germany
An ostracod , a species of Vernoniella .
Viaia [ 74]
Gen. et sp. nov.
Valid
Artal et al.
Cretaceous (Albian or Cenomanian)
Spain
A viaiid glaessneropsoid crab . The type species is Viaia robusta .
Yunnanopalinura [ 64]
Gen. et sp. nov
Valid
Feldmann et al.
Middle or late Anisian
Guanling Formation
China
A decapod crustacean; a spiny lobster . The type species is Yunnanopalinura schrami .
Insects
Blattodea
Coleoptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Abrocar concavus [ 108]
Sp. nov
Valid[ 109]
Davis et al.
Early Cretaceous
Yixian Formation
China
A weevil , a species of Abrocar .
Abrocar relicinus [ 108]
Sp. nov
Valid[ 109]
Davis et al.
Early Cretaceous
Yixian Formation
China
A weevil , a species of Abrocar .
Abrohadeocoleodes [ 110]
Gen. et 4 sp. nov
Valid[ 111]
Tan et al.
Middle Jurassic
Jiulongshan Formation
China
A schizophorid archostematan beetle . Genus contains four species: A. eurycladus , A. ooideus , A. nii and A. patefactus .
Alloioscarabaeus [ 112]
Gen. et sp. nov
Valid
Bai, Ren & Yang in Bai et al.
Middle Jurassic
Jiulongshan Formation
China
An alloioscarabaeid scarabaeoid beetle . The type species is Alloioscarabaeus cheni .
Alphitopsis [ 113]
Gen. et sp. nov
Valid
Kirejtshuk, Nabozhenko & Nel
Early Cretaceous
Yixian Formation
China
A tenebrionine darkling beetle . The type species is Alphitopsis initialis .
Anotylus archaicus [ 114]
sp nov
Valid
Yue, Makranczy & Ren
Early Cretaceous
Yixian Formation
China
An oxyteline rove beetle , a species of Anotylus .
Aphebodactyla [ 115]
Gen. et sp. nov
Valid
Chatzimanolis et al.
Cretaceous
Burmese amber
Myanmar
A ptilodactylid byrrhoid beetle. The type species is Aphebodactyla rhetine .
Aploceble (Chalcoaploceble) viridiaeneus [ 116]
Subgen. et sp. nov
Valid
Tshernyshev
Eocene
Baltic amber
Russia (Kaliningrad Oblast )
A melyrid beetle belonging to the subfamily Dasytinae , a subgenus and species of Aploceble .
Apticax [ 117]
Gen. et 2 sp. nov
Valid
Schomann & Solodovnikov
Early Cretaceous (Aptian–Albian )
Crato Formation
Brazil
A rove beetle of uncertain phylogenetic placement. The type species is A. volans ; genus also includes A. solidus .
Archaeosciaphilus [ 118]
Gen. et sp. nov
Valid
Legalov
Eocene
Europe
A curculionid beetle found in Baltic amber . The type species is Archaeosciaphilus marshalli .
Attagenus yantarnyi [ 119]
Sp. nov
Valid
Háva & Bukejs
Late Eocene
Prussian Formation
Russia (Kaliningrad Oblast )
A dermestid beetle found in Baltic amber , a species of Attagenus .
Baltocar groehni [ 120]
Sp. nov
Valid
Riedel in Riedel et al.
Eocene
Baltic amber
Russia ( Kaliningrad Oblast )
A weevil belonging to the family Attelabidae .
Baltocar hoffeinsorum [ 120]
Sp. nov
Valid
Riedel in Riedel et al.
Eocene
Baltic amber
Russia ( Kaliningrad Oblast )
A weevil belonging to the family Attelabidae.
Baltocar subnudus [ 120]
Sp. nov
Valid
Riedel in Riedel et al.
Eocene
Baltic amber
Russia ( Kaliningrad Oblast )
A weevil belonging to the family Attelabidae.
Cartodere (Aridius) succinobaltica [ 121]
Sp. nov
Valid
Bukejs, Rücker & Kirejtshuk
Late Eocene
Europe
A latridiid beetle found in Baltic amber , a species of Cartodere .
Catops perkovskyi [ 122]
Sp. nov
Valid
Perreau
Eocene
Russia (Kaliningrad Oblast )
A leiodid beetle found in Baltic amber , a species of Catops .
Conapium (Palaeoconapion) alleni [ 123]
Subgen. et sp. nov
Valid
Legalov
Eocene
Baltic amber
Russia (Kaliningrad Oblast )
A beetle belonging to the family Brentidae , a subgenus and species of Conapium .
Crassisorus [ 124]
Gen. et sp. nov
Valid
Nikolajev, Wang & Zhang
Early Cretaceous
Yixian Formation
China
A member of the family Hybosoridae . The type species is C. fractus .
Cretohypna [ 125]
Gen. et sp. nov
Valid
Yan, Nikolajev & Ren
Early Cretaceous
Yixian Formation
China
A glaphyrid beetle . The type species is Cretohypna cristata .
Cretotanaos [ 126]
Gen. et sp. nov
Valid[ 127]
Legalov
Early Cretaceous (Aptian)
Mongolia
An apionine brentid weevil . Type species is Cretotanaos bontsaganensis .
Cryptophagus harenus [ 128]
Sp. nov
Valid
Lyubarsky & Perkovsky
Late Eocene
Ukraine
A cryptophagid beetle found in Rovno amber , a species of Cryptophagus .
Electotreta [ 129]
Gen. et sp.
Valid
Kazantsev
Eocene
Lithuania
A firefly . The type species is Electotreta rasnitsyni .
Electromethes [ 129]
Gen. et sp. nov
Valid
Kazantsev
Eocene
Lithuania
An omethid beetle. The type species is Electromethes alleni .
Electropteron [ 130]
Gen. et sp. nov
Valid
Kazantsev
Oligocene
Dominican Republic
A lycid . The type species is E. avus .
Elodes beigeli [ 131]
Sp. nov
Valid
Klausnitzer
Eocene
Europe
A scirtid beetle found in Baltic amber , a species of Elodes .
Elodes mysticopalpalis [ 131]
Sp. nov
Valid
Klausnitzer
Eocene
Europe
A scirtid beetle found in Baltic amber , a species of Elodes .
Enicmus adrianae [ 132]
Sp. nov
Valid
Bukejs, Reike & Rücker
Late Eocene
Russia (Kaliningrad Oblast )
A member of Latridiidae found in Baltic amber , a species of Enicmus .
Eocenorhynchites [ 118]
Gen. et sp. nov
Valid
Legalov
Eocene
Europe
A rhynchitid beetle found in Baltic amber . The type species is Eocenorhynchites vossi .
Eodromeus daohugouensis [ 133]
sp nov
Valid
Wang, Zhang and Ponomarenko
Middle Jurassic
Daohugou Beds
China
A trachypachid , a species of Eodromeus .
Eodromeus robustus [ 133]
sp nov
Valid
Wang, Zhang and Ponomarenko
Middle Jurassic
Daohugou Beds
China
A trachypachid, a species of Eodromeus .
Eoluciola [ 134]
Gen. et sp. nov
Valid
Kazantsev
Eocene
Europe (Baltic Sea coast)
A lucioline firefly found in Baltic amber . The type species is Eoluciola varang .
Euroleptochromus [ 135]
Gen. et sp. nov
Valid
Jałoszyński
Eocene
Lithuania
A rove beetle found in Lithuanian Eocene Baltic amber . The type species is Euroleptochromus sabathi .
Helophorus (Mesosperchus) inceptivus [ 136]
Sp. nov
Valid
Fikáček et al.
Late Jurassic
Mongolia
A hydrophiloid beetle , a species of Helophorus .
Helophorus (Mesosperchus) yixianus [ 136]
Sp. nov
Valid
Fikáček et al.
Jurassic-Cretaceous boundary
Yixian Formation
China
A hydrophiloid beetle , a species of Helophorus .
Hydroscapha? jeholensis [ 137]
Sp. nov
Valid
Cai, Short & Huang
Early Cretaceous
Yixian Formation
China
A member of the family Hydroscaphidae .
Juroglypholoma [ 138]
Gen. et sp. nov
Valid
Cai et al.
Middle Jurassic
Daohugou Beds
China
A glypholomatine rove beetle . The type species is Juroglypholoma antiquum .
Kararhynchus gratshevi [ 139]
Species
Valid
Legalov
Middle-Late Jurassic
Karabastau Formation
Kazakhstan
An obrieniid beetle , a species of Kararhynchus .
Kararhynchus jurassicus [ 139]
Species
Valid
Legalov
Middle-Late Jurassic
Karabastau Formation
Kazakhstan
An obrieniid beetle, a species of Kararhynchus .
Laetopsia [ 136]
Gen. et comb. nov
Valid
Fikáček et al.
Early Cretaceous
China
Mongolia
Russia
A hydrophiloid beetle ; a new genus for "Hydrophilopsia " baissensis Ponomarenko (1987), "H." bontsaganica Prokin (2009), "H." hydraenoides Prokin, Ren & Fikáček (2010), "H." mongolica Ponomarenko (1987) and "H." shatrovskiyi Prokin, Ren & Fikáček (2010).
Lasiosyne cataphracta [ 140]
Sp. nov
Valid
Yan
Early Cretaceous
Mongolia
A lasiosynid beetle , a species of Lasiosyne .
Lasiosyne decora [ 140]
Sp. nov
Valid
Yan
Early Cretaceous
Russia
A lasiosynid beetle , a species of Lasiosyne .
Lasiosyne insculpta [ 140]
Sp. nov
Valid
Yan
Late Jurassic
Mongolia
A lasiosynid beetle , a species of Lasiosyne .
Lasiosyne longitarsa [ 140]
Sp. nov
Valid
Yan
Late Jurassic
Mongolia
A lasiosynid beetle , a species of Lasiosyne .
Lasiosyne punctata [ 140]
Sp. nov
Valid
Yan
Early Cretaceous
Mongolia
A lasiosynid beetle , a species of Lasiosyne .
Lasiosyne shartegiensis [ 140]
Sp. nov
Valid
Yan
Late Jurassic
Mongolia
A lasiosynid beetle , a species of Lasiosyne .
Leehermania [ 141]
Gen. et sp. nov
Valid
Chatzimanolis, Grimaldi & Engel in Chatzimanolis et al.
Late Triassic (late Carnian or early Norian)
Cow Branch Formation
United States
A beetle of uncertain phylogenetic placement, might be the earliest known rove beetle or a member of Myxophaga related to the family Hydroscaphidae .[ 142] The type species is Leehermania prorova .
Lithohypna lepticephala [ 143]
Sp. nov
Valid
Nikolajev & Ren
Early Cretaceous
Yixian Formation
China
A beetle belonging to the family Glaphyridae , a species of Lithohypna .
Lithohypna longula [ 143]
Sp. nov
Valid
Nikolajev & Ren
Early Cretaceous
Yixian Formation
China
A beetle belonging to the family Glaphyridae , a species of Lithohypna .
Lithohypna tuberculata [ 143]
Sp. nov
Valid
Nikolajev & Ren
Early Cretaceous
Yixian Formation
China
A beetle belonging to the family Glaphyridae , a species of Lithohypna .
Lithohypna yuxiana [ 143]
Sp. nov
Valid
Nikolajev & Ren
Early Cretaceous
Yixian Formation
China
A beetle belonging to the family Glaphyridae , a species of Lithohypna .
Macratria alleni [ 144]
Sp. nov
Valid
Telnov
Late Eocene
Latvia or Lithuania
A macratriine anthicid beetle found in Baltic amber , a species of Macratria .
Melanapion (Melanapionoides) wanati [ 123]
Subgen. et sp. nov
Valid
Legalov
Eocene
Baltic amber
Russia (Kaliningrad Oblast )
A beetle belonging to the family Brentidae , a subgenus and species of Melanapion .
Menopraesagus oryziformis [ 110]
Sp. nov
Valid[ 145]
Tan et al.
Middle Jurassic
Jiulongshan Formation
China
A schizophorid archostematan beetle , a species of Menopraesagus .
Nephus subcircularis [ 146]
Sp. nov
Valid
Kirejtshuk & Nel
Earliest Eocene
France
A scymnine coccinellid beetle found in Oise amber, as species of Nephus
Notocupes lini [ 147]
Sp. nov
Valid
Ponomarenko et al.
Early Jurassic
China
Originally described as a species of Notocupes ; Kirejtshuk (2020) transferred this species to the genus Zygadenia .[ 148]
Olemehlia [ 149]
Gen. et sp. nov
Junior homonym
Batelka
Eocene (Lutetian )
Baltic amber
Lithuania
A member of the family Ripiphoridae belonging to the subfamily Ripidiinae . The type species is O. krali . The generic name is preoccupied by Olemehlia Holzschuh (2011); Batelka (2017) coined a replacement name Olemehliella .[ 150]
Omma delicata [ 151]
Sp. nov
Tan et al.
Middle Jurassic
Jiulongshan Formation
China
A species of Omma
Orchesia (Orchestera) turkini [ 152]
Sp. nov
Valid
Alekseev & Bukejs
Late Eocene
Prussian Formation
Russia (Kaliningrad Oblast )
A melandryid beetle found in Baltic amber , a species of Orchesia .
Palaeoerirhinus [ 126]
Gen. et 4 sp. nov
Valid[ 127]
Legalov
Early Cretaceous (Aptian)
Mongolia
A curculionid weevil . Type species is Palaeoerirhinus ponomarenkoi ; genus also contains new species P. latus , P. thompsoni and P. longirostris .
Palaeometrioxena [ 118]
Gen. et sp. nov
Junior synonym
Legalov
Eocene
Baltic amber
Europe (Baltic Sea region)
A belid . The type species is Palaeometrioxena zherikhini . Legalov (2019) considered the genus Palaeometrioxena to be a junior synonym of the genus Archimetrioxena , and transferred the species P zherikhini to the latter genus.[ 153]
Pareuryomma [ 151]
Nom. et 2 sp. nov
Tan et al.
Early Cretaceous
Yixian Formation
China
A member of the family Ommatidae ; a replacement name for Euryomma Tan et al. (2006). The type species is "Euryomma" tylodes Tan et al. (2006); genus also includes new species P. cardiobasis . Additional new species described by Tan et al. , P. ancistrodonta from the Middle Jurassic Jiulongshan Formation , was subsequently transferred to the genus Omma by Kirejtshuk (2020).[ 148]
Pleocoma dolichophylla [ 154]
Sp. nov
Valid
Nikolajev & Ren
Early Cretaceous
Yixian Formation
China
A rain beetle , a species of Pleocoma .
Prosinodendron [ 155]
Gen. et sp. nov
Valid
Bai, Ren, & Yang
Barremian to early Aptian
Yixian Formation
China
A stag beetle . The type species is Prosinodendron krelli .
Prosolierius [ 156]
Gen. et 3 sp. nov
Valid
Thayer, Newton, & Chatzimanolis
Barremian to Albian
Lebanon
Myanmar
A rove beetle .
Pseudobrienia [ 139]
Gen. et sp. nov
Valid
Legalov
Middle-Late Jurassic
Karabastau Formation
Kazakhstan
An obrieniid beetle. Its type species is Pseudobrienia rasnitsyni .
Pseudoglaesotropis [ 118]
Gen. et sp. nov
Valid
Legalov
Eocene
Baltic amber
Europe (Baltic Sea region)
An anthribid beetle. The type species is Pseudoglaesotropis martynovi . Legalov (2019) subsequently changed the rank of the genus Pseudoglaesotropis to the subgenus of the genus Glaesotropis , and transferred the species P. martynovi to the latter genus.[ 153]
Pulcherhybosorus [ 157]
Gen. et sp. nov
Valid
Yan, Bai & Ren
Early Cretaceous
Yixian Formation
China
A hybosorid beetle. The type species is Pulcherhybosorus tridentatus .
Rhyzobius antiquus [ 146]
Sp. nov
Valid
Kirejtshuk & Nel
Earliest Eocene
Oise amber
France
A cocciduline coccinellid beetle , a species of Rhyzobius
Rhyzobius gratiosus [ 146]
Sp. nov
Valid
Kirejtshuk & Nel
Earliest Eocene
Oise amber
France
A cocciduline coccinellid beetle, a species of Rhyzobius
Septiventer [ 158]
Gen. et sp. nov
Valid[ 159]
Bai et al.
Late Jurassic or Early Cretaceous
Yixian Formation
China
A septiventerid scarabaeoid beetle . The type species is Septiventer quadridentatus .
Shartegopsis [ 160]
Gen. et sp. nov
Valid
Kirejtshuk
Late Jurassic
Mongolia
A clerid beetle . The type species is Shartegopsis miranda .
Sinodromeus [ 133]
Gen. et sp. nov
Valid
Wang, Zhang & Ponomarenko
Early Cretaceous
Yixian Formation
China
A trachypachid. The type species is Sinodromeus liutiaogouensis .
Sinopeltis [ 161]
Gen. et 2 sp. nov
Valid
Yu et al.
Middle Jurassic
Jiulongshan Formation
China
A peltine trogossitid beetle . Genus contains two species: S. jurrasica (type species) and S. amoena . Kirejtshuk (2017) considered the genus Sinopeltis to be a junior synonym of the genus Cervicatinius Tan & Ren (2007).[ 162]
Sinoschizala [ 163]
Gen. et sp. nov
Valid
Jarzembowski et al.
Middle Jurassic
Daohugou Beds
China
A schizophorid beetle . The type species is Sinoschizala darani .
Succinometrioxena [ 164]
Gen. et sp. nov
Valid
Legalov
Eocene
Europe
A belid weevil found in Baltic amber . The type species is Succinometrioxena poinari .
Sucinoptinus bukejsi [ 165]
Sp. nov
Valid
Alekseev
Eocene (Priabonian )
Prussian Formation
Russia (Kaliningrad Oblast )
A spider beetle found in Baltic amber , a species of Sucinoptinus .
Tafforeus [ 122]
Gen. et sp. nov
Valid
Perreau
Eocene
Russia (Kaliningrad Oblast )
A leiodid beetle found in Baltic amber . The type species is Tafforeus cainosternus .
Talbragarus [ 166]
Gen. et sp. nov
Valid
Oberprieler & Oberprieler
Late Jurassic
Talbragar Fish Bed
Australia
A nemonychid weevil . The type species is Talbragarus averyi .
Tetraphalerus decorosus [ 151]
Sp. nov
Tan et al.
Middle Jurassic
Jiulongshan Formation
China
Originally described as a species of Tetraphalerus ; Kirejtshuk (2020) transferred this species to the ommatine genus Monticupes .[ 148]
Toksunius [ 147]
Gen. et comb. nov
Valid
Ponomarenko et al.
Late Triassic
China
A new genus for "Ademosynoides " chinaticus Lin (1992).
Tomoderus balticus [ 167]
Sp. nov
Valid
Telnov
Late Eocene
Latvia or Lithuania
A tomoderine anthicid beetle found in Baltic amber , a species of Tomoderus .
Trixagus majusculus [ 168]
sp nov
Valid
Kovalev, Kirejtshuk & Nel
Lowermost Eocene
France
A throscid beetle , a species of Trixagus .
Unda chifengensis [ 133]
sp nov
Valid
Wang, Zhang & Ponomarenko
Middle Jurassic
Daohugou Beds
China
A trachypachid, a species of Unda .
Xenohimatium [ 169]
Gen. et sp. nov
Valid
Lyubarsky & Perkovsky
Late Eocene
Rovno amber
Ukraine
A pleasing fungus beetle belonging to the subfamily Xenoscelinae . The type species is X. rovnense .
Diptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Aetheapnomyia [ 170]
Gen. et comb. nov
Valid
Harbach & Greenwalt
Eocene
Europe
A mosquito found in Baltic Amber , a new genus for "Aedes " hoffeinsorum Szadziewski 1998.
Bibio anatolicus [ 171]
Sp. nov
Valid
Skartveit & Nel
Oligocene
Turkey
A species of Bibio .
Bibio nigricosta [ 171]
Sp. nov
Valid
Skartveit & Nel
Oligocene
Turkey
A species of Bibio .
Calosargus talbragarensis [ 172]
Sp. nov
Valid
Oberprieler & Yeates
Late Jurassic
Australia
An archisargid brachyceran , a species of Calosargus .
Clarumreddera [ 173]
Gen. et 2 sp. nov
Valid
Fedotova & Perkovsky
Late Eocene
Rovno amber
Ukraine
A cecidomyiid fly , with two species: C. korneyevi and C. conceptiva .
Culiseta kishenehn [ 170]
Sp. nov
Valid
Harbach & Greenwalt
Eocene
Kishenehn Formation
United States
A mosquito , a species of Culiseta .
Culiseta lemniscata [ 170]
Sp. nov
Valid
Harbach & Greenwalt
Eocene
Kishenehn Formation
United States
A mosquito , a species of Culiseta .
Curtonotum electrodominicum [ 174]
Sp. nov
Valid
Grimaldi & Kirk-Spriggs
Miocene
Dominican Republic
A curtonotid found in Dominican amber , a species of Curtonotum .
Daohugosargus [ 175]
Gen. et comb. nov
Valid
Zhang
Middle Jurassic
Daohugou Beds
China
An archisargid brachyceran ; a new genus for "Sharasargus " eximius Zhang, Yang & Ren (2008).
Depressonotum [ 174]
Gen. et sp. nov
Valid
Grimaldi & Kirk-Spriggs
Miocene
Dominican Republic
A curtonotid found in Dominican amber . The type species is Depressonotum priscum .
Dicranomyia (Caenolimonia) alexbrowni [ 176]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Dicranomyia .
Dicranomyia (Dicranomyia) chiapa [ 176]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Dicranomyia .
Dicranomyia (Dicranomyia) mexa [ 176]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Dicranomyia .
Dicranomyia (Dicranomyia) vella [ 176]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Dicranomyia .
Didactylomyia dlusskyi [ 177]
Sp. nov
Valid
Fedotova & Perkovsky
Late Eocene
Ukraine
A cecidomyiid fly found in Rovno amber, a species of Didactylomyia .
Enischnomyia [ 178]
Gen. et sp. nov
Valid
Poinar & Brown
Burdigalian
Dominican amber
Dominican Republic
A bat fly, type species E. stegosoma , host for the Plasmodiidae Vetufebrus ovatus [ 179]
Enischnomyia stegosoma
Eoaedes [ 170]
Gen. et comb. nov
Valid
Harbach & Greenwalt
Eocene
Europe
A mosquito found in Baltic Amber , a new genus for "Aedes " damzeni Szadziewski 1998.
Eoptychopterina postica [ 180]
Sp. nov
Valid
Liu, Shih & Ren
Middle Jurassic
Jiulongshan Formation
China
A ptychopterid , a species of Eoptychopterina .
Eotrichocera (Archaeotrichocera) spatiosa [ 180]
Sp. nov
Valid
Liu, Shih & Ren
Middle Jurassic
Jiulongshan Formation
China
A trichocerid , a species of Eotrichocera .
Erioptera (?Mesocyphona) beata [ 176]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Erioptera .
Erioptera (Mesocyphona) diver [ 176]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Erioptera .
Erioptera (?Mesocyphona) sherry [ 176]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Erioptera .
Flagellisargus [ 181]
Gen. et 3 sp. nov
Valid
Zhang
Callovian or Oxfordian
Daohugou Beds
China
An archisargid fly. Genus contains three species: Flagellisargus sinicus , F. venustus and F. robustus .
Groveromyia [ 182]
Gen. et 3 sp. et comb. nov
Valid
Fedotova & Perkovsky
Late Eocene
Rovno amber
Ukraine
A cecidomyiid fly with 4 species G. occlusa ; G. digna , G. astrosa , and "Bryocrypta " concinna Fedotova (2005).
Helius (Helius) collemus [ 176]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Helius .
Jurochlus adustus [ 183]
Sp. nov
Valid
Lukashevich & Przhiboro
Mesozoic
Mongolia
A chironomid , a species of Jurochlus .
Jurochlus limbatus [ 183]
Sp. nov
Valid
Lukashevich & Przhiboro
Mesozoic
Mongolia
A chironomid , a species of Jurochlus .
Jurochlus lineatus [ 183]
Sp. nov
Valid
Lukashevich & Przhiboro
Mesozoic
Mongolia
A chironomid , a species of Jurochlus .
Jurochlus trivittatus [ 183]
Sp. nov
Valid
Lukashevich & Przhiboro
Mesozoic
Mongolia
A chironomid , a species of Jurochlus .
Lacrimyza [ 184]
Gen. et 2 sp. nov
Valid[ 185]
Roháček
Eocene
Europe
An anthomyzid . Genus contains two species: L. lacrimosa and L. christelae .
Laiyangitabanus [ 186]
Gen. et sp. nov
Valid
Zhang
Early Cretaceous
Laiyang Formation
China
A horse-fly . The type species is Laiyangitabanus formosus .
Lithorhagio [ 187]
Gen. et sp. nov
Valid
Zhang & Li
Middle Jurassic or lowermost Upper Jurassic
Daohugou Beds
China
A rhagionid . The type species is Lithorhagio megalocephalus .
Lygistorrhina caribbiana [ 188]
Sp. nov
Valid
Grund
Miocene
Dominican Republic
A lygistorrhinid fungus gnat found in Dominican amber , a species of Lygistorrhina .
Mesorhyphus handlirschi [ 189]
Sp. nov
Valid
Lukashevich
Upper Jurassic
Mongolia
An anisopodid bibionomorph , a species of Mesorhyphus .
Mesorhyphus hennigi [ 189]
Sp. nov
Valid
Lukashevich
Upper Jurassic
Mongolia
An anisopodid bibionomorph , a species of Mesorhyphus .
Mesosciophila sigmoidea [ 190]
sp nov
Valid
Wang, Zhao & Ren
Middle Jurassic
Jiulongshan Formation
China
A mesosciophilid nematoceran , a species of Mesosciophila .
Mesosolva zhangae [ 175]
Nom. nov
Valid
Zhang
Middle Jurassic
Daohugou Beds
China
An archisargid brachyceran ; a replacement name for Brevisolva daohugouensis Zhang et al. (2010).
Nemopalpus inexpectatus [ 191]
Sp. nov
Valid
Wagner in Wagner & Stuckenberg
Late Eocene
Baltic amber
Europe
A bruchomyiine psychodid fly . Originally described as a species of Nemopalpus , but subsequently transferred to the genus Palaeosycorax .[ 192]
Nemopalpus velteni [ 191]
Sp. nov
Valid
Wagner in Wagner & Stuckenberg
Cretaceous (Albian)
Burmese amber
Myanmar
A bruchomyiine psychodid fly . Originally described as a species of Nemopalpus , but subsequently transferred to the genus Palaeoglaesum .[ 192]
Orientisargus [ 175]
Gen. et sp. nov
Valid
Zhang
Latest Middle Jurassic or earliest Late Jurassic
Daohugou Beds
China
An orientisargid archisargoid brachyceran . The type species is Orientisargus illecebrosus .
Oryctochlus brundini [ 193]
Sp. nov
Valid
Lukashevich
Late Jurassic
Mongolia
A podonomine chironomid , as species of Oryctochlus
Oryctochlus kaluginae [ 193]
Sp. nov
Valid
Lukashevich
Late Jurassic
Mongolia
A podonomine chironomid , as species of Oryctochlus
Palaeopoecilostola eocenica [ 194]
Sp. nov
Valid
Kania & Krzemiński
Eocene
Europe
A limoniid found in Baltic amber , a species of Palaeopoecilostola .
Paramesosciophilodes aequus [ 190]
sp nov
Valid
Wang, Zhao & Ren
Middle Jurassic
Jiulongshan Formation
China
A mesosciophilid nematoceran , a species of Paramesosciophilodes .
Penthetria beskonakensis [ 171]
Sp. nov
Valid
Skartveit & Nel
Oligocene
Turkey
A species of Penthetria .
Probittacomorpha brisaci [ 195]
sp nov
Valid
Krzemiński, Kania & Nel
Late Miocene
France
A ptychopterid , a species of Probittacomorpha .
Procramptonomyia kovalevi [ 189]
Sp. nov
Valid
Lukashevich
Upper Jurassic
Kazakhstan
Mongolia
A procramptonomyiid bibionomorph , a species of Procramptonomyia .
Procramptonomyia ponomarenkoi [ 189]
Sp. nov
Valid
Lukashevich
Upper Jurassic
Mongolia
A procramptonomyiid bibionomorph , a species of Procramptonomyia .
Protanthomyza hennigi [ 184]
Sp. nov
Valid[ 185]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protanthomyza hoffeinsorum [ 184]
Sp. nov
Valid[ 185]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protanthomyza krylovi [ 184]
Sp. nov
Valid[ 185]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protanthomyza loewi [ 184]
Sp. nov
Valid[ 185]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protanthomyza meunieri [ 184]
Sp. nov
Valid[ 185]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protanthomyza presli [ 184]
Sp. nov
Valid[ 185]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protanthomyza tschirnhausi [ 184]
Sp. nov
Valid[ 185]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protoculicoides ciliatus [ 196]
Sp. nov
Valid
Borkent
Cretaceous
Canadian amber
Canada
A member of the family Ceratopogonidae . Originally described as a species of Protoculicoides ; Borkent (2019) transferred this species to the genus Atriculicoides .[ 197]
Protorhagio parvus [ 187]
sp nov
Valid
Zhang & Li
Middle Jurassic or lowermost Upper Jurassic
Daohugou Beds
China
A rhagionid, a species of Protorhagio .
Protorhyphus rohdendorfi [ 189]
Sp. nov
Valid
Lukashevich
Upper Jurassic
Mongolia
A protorhyphid bibionomorph , a species of Protorhyphus .
Rovnodidactylomyia [ 177]
Gen. et 3 sp. et comb. nov
Valid
Fedotova & Perkovsky
Late Eocene
Europe
A cecidomyiid fly with 5 species R. zosimovichi , R. sidorenkoi , R. iconica ) in Rovno amber, plus "Bryocrypta " girafa Meunier (1904) and "B." capitosa Meunier (1904) in Baltic amber .
Sinallomyia [ 175]
Nom. nov
Valid
Zhang
Early Cretaceous
Yixian Formation
China
An archisargid brachyceran ; a replacement name for Allomyia Ren (1998).
Sinocretomyia [ 186]
Gen. et sp. nov
Valid
Zhang
Early Cretaceous
Laiyang Formation
China
An athericid . The type species is Sinocretomyia minuscula .
Stilobezzia pikei [ 196]
Sp. nov
Valid
Borkent
Cretaceous
Canadian amber
Canada
A species of Stilobezzia .
Stomatosema iljieteugeniae [ 173]
Sp. nov
Valid
Fedotova & Perkovsky
Late Eocene
Rovno amber
Ukraine
A cecidomyiid fly
Tipula paleopannonia [ 198]
Sp. nov
Valid
Engel & Gross
Miocene (ca. 11.3 Ma)
Paldau Formation
Austria
A crane fly , a species of Tipula .
Ephemeroptera
Hemiptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Acaricoris robertae [ 203]
Sp. nov
Valid
Heiss & Poinar
Miocene
Dominican Republic
An aradid found in Dominican amber , a species of Acaricoris .
Arctorthezia baltica [ 204]
Sp. nov
Valid
Vea & Grimaldi
Eocene
Europe
An ortheziid found in Eocene Baltic amber , a species of Arctorthezia .
Breviscutum [ 205]
Gen. et sp. nov
Valid[ 206]
Yao et al.
Cretaceous
China
A primipentatomid pentatomoid . The type species is Breviscutum lunatum .
Burmacicada [ 207]
Gen. et sp. nov
Valid
Poinar & Kritsky
Cretaceous
Burmese amber
Myanmar
A cicada . The type species is Burmacicada protera .
Burmorthezia [ 204]
Gen et 2 sp. nov
Valid
Vea & Grimaldi
Cretaceous (98 Ma)
Burmese amber
Myanmar
An ortheziid . Genus contains two species: Burmorthezia kotejai and B. insolita .
Carventus bechlyi [ 203]
Sp. nov
Valid
Heiss & Poinar
Miocene
Dominican Republic
An aradid found in Dominican amber , a species of Carventus .
Cicadocoris varians [ 208]
Sp. nov
Disputed
Dong, Yao & Ren
Middle Jurassic
Jiulongshan Formation
China
A moss bug belonging to the family Progonocimicidae . Jiang & Huang (2017) considered this species to be a probable junior synonym of Cicadocoris sinensis Hong (1983).[ 209]
Cinemala [ 210]
Gen. et sp. nov
Valid
Shcherbakov
Middle or Late Jurassic
Bakhar Group
Mongolia
A member of Cicadomorpha belonging to the family Hylicellidae . The type species is C. contrasta .
Cintux [ 211]
Gen. et sp. nov
Valid
Stroiński & Szwedo
Palaeocene
France
A lophopid planthopper . The type species is Cintux menatensis .
Conjucella [ 210]
Gen. et sp. nov
Valid
Shcherbakov
Early Cretaceous
Zaza Formation
Russia
A member of Cicadomorpha belonging to the family Hylicellidae . The type species is C. selene .
Dellashara [ 212]
Gen. et sp. nov
Valid
Shcherbakov
Late Jurassic
Shar Teg Sequence
Mongolia
A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae . The type species is D. tega .
Dominicicada [ 207]
Gen. et sp. nov
Valid
Poinar & Kritsky
Dominican Republic
A cicada found in Dominican amber . The type species is Dominicicada youngi .
Edessa protera [ 213]
Sp. nov
Valid
Poinar & Thomas
Tertiary
Mexico
A pentatomid stink bug found in Mexican amber, a species of Edessa .
Hispanocader [ 214]
Gen. et sp. nov
Valid
Golub, Popov & Arillo
Albian
Escucha Formation
Spain
A relative of tingids . The type species is Hispanocader lisae .
Hoffeinsoria [ 215]
Gen. et sp. nov
Valid
Herczek & Popov
Eocene
Prussian Formation
Europe
An isometopine mirid found in Eocene Baltic amber . The type species is Hoffeinsoria robusta .
Jiania [ 216]
Gen. et 2 sp. nov
Valid
Wang & Szwedo in Wang, Szwedo & Zhang
Middle Jurassic
Daohugou Beds
China
A sinoalid froghopper . Genus contains two species: Jiania crebra and Jiania gracila . Fu, Cai & Huang (2018) considered J. gracila to be a junior synonym of J crebra .[ 217]
Kachinocoris [ 218]
Gen. et sp. nov
Valid
Heiss
Cretaceous
Burmese amber
Myanmar
A flat bug found in Cretaceous Burmese Amber. The type species is Kachinocoris brevipennis .
Kemobius [ 212]
Gen. et sp. nov
Valid
Shcherbakov
Middle Jurassic
Itat Formation
Russia
A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae . The type species is K. lux .
Kisa [ 212]
Gen. et sp. nov
Valid
Shcherbakov
Middle Jurassic
Itat Formation
Russia
A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae . The type species is K. fasciata .
Kubecola [ 212]
Gen. et sp. nov
Valid
Shcherbakov
Middle Jurassic
Itat Formation
Russia
A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae . The type species is K. guttatus .
Microaradus [ 219]
Gen. et sp. nov
Valid
Heiss & Poinar
Mesozoic
Burmese amber
Myanmar
An archaearadine aradid found in Mesozoic Burmese amber. The type species is Microaradus anticus .
Minyscapheus [ 220]
Gen. et sp. nov.
Valid
Poinar, Kritsky & Brown
Dominican Republic
A cicada found in Dominican amber . The type species is Minyscapheus dominicanus .
Mixorthezia dominicana [ 204]
Sp. nov
Valid
Vea & Grimaldi
Miocene
Dominican Republic
An ortheziid found in Miocene Dominican amber , a species of Mixorthezia .
Mixorthezia kozari [ 204]
Sp. nov
Valid
Vea & Grimaldi
Miocene
Dominican Republic
An ortheziid found in Miocene Dominican amber , a species of Mixorthezia .
Mongolocoris [ 221]
Gen. et comb. nov
Valid
Ryzhkova
Early Cretaceous
Mongolia
An enicocorid leptopodomorph heteropteran , a new genus for "Enicocoris " tibialis Popov, 1986.
Myanmezira [ 219]
Gen. et sp. nov
Valid
Heiss & Poinar
Mesozoic
Burmese amber
Myanmar
A mezirine aradid found in Mesozoic Burmese amber. The type species is Myanmezira longicornis .
Nesoproxius latocanus [ 203]
Sp. nov
Valid
Heiss & Poinar
Miocene
Dominican Republic
An aradid found in Dominican amber , a species of Nesoproxius .
Notonecta vetula [ 222]
sp nov
Valid
Zhang, Yao & Ren
Late Jurassic
Chijinqiao Formation
China
A notonectid ,a species of Notonecta .
Oropentatoma [ 205]
Gen. et sp. nov
Valid[ 223]
Yao et al.
Cretaceous
China
A primipentatomid pentatomoid . The type species is Oropentatoma epichara .
Poljanka hirsuta [ 224]
sp nov
Valid
Yang, Yao & Ren
Middle Jurassic
Jiulongshan Formation
China
A member of Sternorrhyncha , a protopsyllidiid ; a species of Poljanka .
Primipentatoma [ 205]
Gen. et 2 sp. nov
Valid[ 225]
Yao et al.
Cretaceous
China
A primipentatomid pentatomoid . Genus contains two species: Primipentatoma peregrina and Primipentatoma fangi .
Proneoproxius [ 203]
Gen. et sp. nov
Valid
Heiss & Poinar
Miocene
Dominican Republic
An aradid found in Dominican amber . The type species is Proneoproxius cornutus .
Purbecellus [ 210]
Gen. et comb. nov
Valid
Shcherbakov
Early Cretaceous
Purbeck Group
United Kingdom
A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae . The type species is "Cicadellium " psocus Westwood (1854).
Quadrocoris [ 205]
Gen. et sp. nov
Valid[ 226]
Yao et al.
Cretaceous
China
A primipentatomid pentatomoid . The type species is Quadrocoris radius .
Sinaldocader rasnitsyni [ 227]
Sp. nov
Valid
Golub & Popov
Turonian
Kazakhstan
A lace bug . Originally described as a species of Sinaldocader ; Popov & Golub (2019) transferred this species to the separate genus Kzylcader .[ 228]
Sinoala [ 216]
Gen. et sp. nov
Valid
Wang & Szwedo in Wang, Szwedo & Zhang
Middle Jurassic
Daohugou Beds
China
A sinoalid froghopper . The type species is Sinoala parallelivena .
Sunotettigarcta hirsuta [ 229]
sp nov
Valid
Li, Wang & Ren in Li et al.
Middle Jurassic
Daohugou Beds
China
A tettigarctid , a species of Sunotettigarcta .
Ulanocoris [ 221]
Gen. et 2 sp. nov
Valid
Ryzhkova
Early Cretaceous
Mongolia
An enicocorid leptopodomorph heteropteran . Genus contains two species: Ulanocoris femoralis and U. grandis .
Venustsalda [ 230]
Gen. et sp. nov
Valid
Zhang et al.
Lower Cretaceous
Yixian Formation
China
A saldid heteropteran . The type species is Venustsalda locella .
Hymenoptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Agapia [ 231]
Gen. et sp. nov
Valid
Kopylov
Late Cretaceous
Russia
An ichneumonid . The type species is Agapia sukatchevae .
Agapteron [ 231]
Gen. et sp. nov
Valid
Kopylov
Late Cretaceous
Russia
An ichneumonid . The type species is Agapteron popovi .
Archimyrmex wedmannae [ 232]
Sp. nov
Valid
Dlussky
Middle Eocene
Messel Pit
Germany
A myrmeciine ant .
Archimyrmex wedmannae
Anthophorula (Anthophorula) persephone [ 233]
Sp. nov
Valid
Engel in Engel et al.
Early Miocene
Dominican Republic
An exomalopsine apid bee found in Dominican amber , a species of Anthophorula .
Atefia [ 234]
Gen. et sp. nov
Valid[ 235]
Krogmann et al.
Early Cretaceous
Crato Formation
Brazil
A tenthredinoid hymenopteran sharing morphological characters with the families Pergidae and Diprionidae . The type species is Atefia rasnitsyni .
Boltonidris [ 236]
Gen. et sp. nov
Valid
Radchenko & Dlussky
Late Eocene
Rovno amber
Ukraine
A stenammine myrmicine ant found in Rovno amber. The type species is Boltonidris mirabilis .
Bombus (Bombus) randeckensis [ 237]
Sp. nov
Valid
Wappler & Engel in Wappler et al.
Miocene
Germany
A bumblebee .
Casaleia eocenica [ 238] [ 239]
Sp nov
valid
Dlussky & Wedmann
Lutetian
Messel Formation
Germany
An amblyoponine ant.
Casaleia eocenica
Cascoscelio [ 5]
Gen. et sp. nov.
Valid
Poinar in Poinar & Buckley
Early Cretaceous
Burmese amber
Myanmar
A platygastrid wasp, type species is C. incassus .
Cascoscelio incassa
Cephalopone [ 238]
Gen. et 2 sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A poneromorph ant . Two species: C. potens & C. grandis .
Cephalopone grandis
Cratoenigma [ 240]
Gen. et sp. nov
Valid
Krogmann & Nél
Early Cretaceous (Aptian )
Crato Formation
Brazil
A probable relative of xiphydriids and euhymenopterans . The type species is C. articulata .
Cretevania soplaensis [ 241]
Sp. nov
Valid
De La Fuente, Peñalver & Ortega-Blanco
Albian
Spain
An evaniid , a species of Cretevania .
Cretorhyssalus [ 242]
Gen. et sp. nov
Valid
Belokobylskij
Cenomanian
Ola Formation
Russia
A braconid wasp . Type species is Cretorhyssalus brevis .
Cyrtopone [ 238]
Gen. et 4 sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel Formation
Germany
A ponerinae ant . Genus contains four species: Cyrtopone curiosa , C. elongata , C. microcephala and C. striata .
C. striata
Gulgonga [ 243]
Gen. et sp. nov
Valid
Oberprieler, Rasnitsyn & Brothers
Late Jurassic
Talbragar Fish Bed
Australia
A praeaulacid evanioid . The type species is Gulgonga beattiei .
Haidomyrmex scimitarus [ 244]
Sp. nov
Valid
Barden & Grimaldi
Late Aptian to Early Cenomanian
Burmese amber
Myanmar
A Sphecomyrmin ant
Haidomyrmex scimitarus
Haidomyrmex zigrasi [ 244]
Sp. nov
Valid
Barden & Grimaldi
Campanian
Burmese amber
Myanmar
A Sphecomyrmin ant
Haidomyrmex zigrasi
Hyptia deansi [ 245]
Sp. nov
Valid
Jennings, Krogmann & Mew
Late Oligocene or early Miocene
Mexico
An evaniid hymenopteran , a species of Hyptia .
Ibalia (Ibalia) electra [ 246]
Sp. nov
Valid
Engel & Liu
Eocene
Europe
An ibaliid found in Baltic amber , a species of Ibalia .
Lancepyris [ 247]
Gen. et sp. nov
Valid
Azevedo & Azar
Early Cretaceous
Lebanon
A lancepyrine bethylid found in Early Cretaceous Lebanese amber. The type species is Lancepyris opertus .
Lonchodryinus balticus [ 248]
Sp. nov
Valid
Olmi & Guglielmino
Late Eocene
Europe
A dryinid found in Baltic amber , a species of Lonchodryinus .
Magadanobracon [ 242]
Gen. et 2 sp. nov
Valid
Belokobylskij
Cenomanian
Ola Formation
Russia
A braconid wasp. The type species is M. rasnitsyni ; a second species is M. zherikhini .
Messelepone [ 238]
Gen. et sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A poneromorph ant . The type species is M. leptogenoides .
Messelepone leptogenoides
Mirabythus [ 249]
Gen. et 2 sp. nov
Valid
Cai, Shih & Ren in Cai et al.
Early Cretaceous
Yixian Formation
China
A scolebythid chrysidoid . The genus contains two species: Mirabythus lechrius and M. liae .
Pachycondyla eocenica [ 238]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A Ponerinae ant .
Pachycondyla eocenica
Pachycondyla lutzi [ 238]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant
Pachycondyla lutzi
Pachycondyla? messeliana [ 238]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant .
Pachycondyla? messeliana
Pachycondyla minuta [ 238] [ 250]
Sp. nov
synonym
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant , name occupied, renamed P. parvula .
Pachycondyla parvula
Pachycondyla petiolosa [ 238]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant .
Pachycondyla petiolosa
Pachycondyla petrosa [ 238]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A poneromorph ant , a species of Pachycondyla .
Pachycondyla petrosa
Platyxyela [ 251]
Gen. et sp. nov
Valid
Wang, Shih & Ren
Middle Jurassic
Jiulongshan Formation
China
A macroxyeline xyelid sawfly . The type species is Platyxyela unica .
Praeichneumon dzhidensis [ 252]
sp nov
Valid
Kopylov
Lower Cretaceous
Russia
A praeichneumonid ichneumonoid , a species of Praeichneumon .
Praeichneumon khamardabanicus [ 252]
sp nov
Valid
Kopylov
Lower Cretaceous
Russia
A praeichneumonid ichneumonoid, a species of Praeichneumon .
Praeichneumon zakhaaminicus [ 252]
sp nov
Valid
Kopylov
Lower Cretaceous
Russia
A praeichneumonid ichneumonoid, a species of Praeichneumon .
Prionomyrmex wappleri [ 232]
Sp. nov
Valid
Dlussky
Late Oligocene
Germany
A myrmeciine ant .
Prionomyrmex wappleri
Protodryinus [ 248]
Gen. et sp. nov
Valid
Olmi & Guglielmino
Late Eocene
Europe
A dryinid found in Baltic amber . The type species is Protodryinus eocenicus .
Protopone? dubia [ 238]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel Formation
Germany
A ponerine ant
Protopone? dubia
Protopone germanica [ 238]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant
Protopone germanica
Protopone magna [ 238]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant
Protopone magna
Protopone oculata [ 238]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant
Protopone oculata
Protopone sepulta [ 238]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant
Protopone sepulta
Protopone vetula [ 238]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant
Protopone vetula
Pseudectatomma [ 238]
Gen. et 2 sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
An Ectatommin ant . Two species: P. eocenica and P. striatula .
Pseudectatomma eocenica
Trichelyon [ 253]
Gen. et sp. nov
Disputed
Ortega-Blanco & Engel in Ortega-Blanco, Singh & Engel
Early Eocene (Ypresian )
India
A rhysipoline exothecine braconid wasp found in Cambay amber. The type species is Trichelyon tadkeshwarense . Belokobylskij & Manukyan (2024) considered Trichelyon to be a junior synonym of the genus Clinocentrus , though the authors maintained T. tadkeshwarense as a distinct species within the latter genus, resulting in a new combination Clinocentrus tadkeshwarense .[ 254]
Urotryphon baikurensis [ 231]
Sp. nov
Valid
Kopylov
Late Cretaceous
Russia
An ichneumonid , a species of Urotryphon .
Zatania electra [ 255]
Sp. nov
Valid
Lapolla, Kallal & Brady
Eocene or Miocene
Dominican amber
Dominican Republic
A formicine ant , a species of Zatania .
Zatania electra
Mecoptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Baltipanorpa [ 256]
Gen. et sp. nov
Valid
Krzemiński & Soszyńska-Maj
Middle Eocene
Baltic amber
Poland
A panorpid scorpionfly . The type species is B. damzeni .
Exilibittacus [ 257]
Gen. et sp. nov
Valid
Yang, Ren & Shih
Middle Jurassic
Jiulongshan Formation
China
A bittacid hangingfly . The type species is E. lii .
Exilibittacus lii
Jeholopsyche bella [ 258]
Sp. nov
Valid
Qiao, Shih & Ren
Early Cretaceous
Yixian Formation
China
An aneuretopsychid mecopteran .
Jeholopsyche completa [ 258]
Sp. nov
Valid
Qiao, Shih & Ren
Early Cretaceous
Yixian Formation
China
An aneuretopsychid mecopteran .
Jeholopsyche maxima [ 258]
Sp. nov
Valid
Qiao, Shih & Ren
Early Cretaceous
Yixian Formation
China
An aneuretopsychid mecopteran .
Juracimbrophlebia [ 259]
Gen. et sp. nov
Valid
Wang, Labandeira, Shih & Ren in Wang et al.
Middle Jurassic (Bathonian or Callovian)
Jiulongshan Formation
China
A cimbrophlebiid mecopteran . Adults with Yimaia ginkglaean leaf mimicry . The type species is J. ginkgofolia .
Juracimbrophlebia ginkgofolia
Megabittacus spatiosus [ 257]
Sp. nov
Valid
Yang, Ren & Shih
Late Jurassic or Early Cretaceous
Yixian Formation
China
A hangingfly , a species of Megabittacus .
Orthophlebia nervulosa [ 260]
sp nov
Valid
Qiao, Shih & Ren
Middle Jurassic
Jiulongshan Formation
China
A mecopteran , a species of Orthophlebia .
Orthophlebia riccardii [ 261]
Sp. nov
Valid
Petrulevičius & Ren
Middle Jurassic
Jiulongshan Formation
China
A mecopteran . Originally described as a species of Orthophlebia , subsequently transferred to the separate genus Gigaphlebia .[ 262]
Orthophlebia stigmosa [ 260]
sp nov
Valid
Qiao, Shih & Ren
Middle Jurassic
Jiulongshan Formation
China
A mecopteran . Originally described as a species of Orthophlebia , subsequently transferred to the separate genus Longiphlebia .[ 262]
Perfecticimbrophlebia [ 263]
Gen. et sp. nov
Valid
Yang, Shih & Ren in Yang et al.
Middle Jurassic
Jiulongshan Formation
China
A cimbrophlebiid mecopteran . The type species is Perfecticimbrophlebia laetus .
Preanabittacus validus [ 263]
sp nov
Valid
Yang, Shih & Ren in Yang et al.
Middle Jurassic
Jiulongshan Formation
China
A hangingfly , a species of Preanabittacus .
Neuroptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Alloepipsychopsis [ 264]
Gen. et sp. nov
Valid
Makarkin, Yang, et al.
Barremian
Yixian Formation
China
A psychopsid . Type species is Alloepipsychopsis lata .
Chorilingia [ 265]
Gen. et 4 sp. nov
Valid
Shi et al.
Middle Jurassic
Jiulongshan Formation
China
A grammolingiid neuropteran . Genus contains four species: of C. euryptera , C. parvica , C. translucida and C. peregrina .
Guyiling [ 266]
Gen. et sp. nov
Valid
Shi et al.
Early Cretaceous
Yixian Formation
China
A member of Myrmeleontiformia of uncertain phylogenetic placement. The type species is G. jianboni .
Hallucinochrysa [ 267]
Gen. et sp. nov
Valid
Pérez-de la Fuente, Delclòs, Peñalver & Engel in Pérez-de la Fuente et al.
Early Cretaceous (Albian)
Spain
A chrysopoid neuropteran (a relative of the green lacewings ). The type species is Hallucinochrysa diogenesi .
Kareninoides [ 268]
Gen. et sp. et comb. nov
Valid
Yang, Makarkin & Ren
Early Cretaceous
La Huerguina Formation
Yixian Formation
China
Spain
A mesochrysopid neuropteran . The type species is Kareninoides lii ; genus might also contain "Armandochrysopa" inexpecta Nel et al. (2005).
Leptolingia oblonga [ 269]
Sp. nov
Valid
Khramov
Late Jurassic
Mongolia
A grammolingiid neuropteran , a species of Leptolingia .
Litholingia longa [ 269]
Sp. nov
Valid
Khramov
Early or Middle Jurassic
Kyrgyzstan
A grammolingiid neuropteran , a species of Litholingia .
Parakseneura [ 270]
Gen. et 11 sp. nov
Valid
Yang, Makarkin & Ren in Yang et al.
Middle Jurassic
Jiulongshan Formation
China
A parakseneurid neuropteran . Genus contains 11 species.
Protoaristenymphes daohugouensis [ 268]
Sp. nov
Valid
Yang, Makarkin & Ren
Middle Jurassic
Daohugou Beds
China
A mesochrysopid neuropteran , a species of Protoaristenymphes .
Protolingia [ 269]
Gen. et sp. nov
Valid
Khramov
Early or Middle Jurassic
Kyrgyzstan
A grammolingiid neuropteran . The type species is Protolingia mira .
Pseudorapisma [ 270]
Gen. et 3 sp. nov
Valid
Yang, Makarkin & Ren in Yang et al.
Middle Jurassic
Jiulongshan Formation
China
A parakseneurid neuropteran . Genus contains 3 species: P. jurassicum , P. maculatum and P. angustipenne .
Shuraboneura [ 270]
Gen. et sp. nov
Valid
Khramov & Makarkin in Yang et al.
Early or Middle Jurassic
Sogul Formation
Kyrgyzstan
A parakseneurid neuropteran . The type species is Shuraboneura ovata .
Notoptera
Odonatoptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Austroprotolindenia [ 277]
Gen. et sp. nov
Valid
Beattie & Nel
Late Jurassic
Talbragar Fossil Fish Bed
Australia
A dragonfly . The type species is Austroprotolindenia jurassica .
Azaroneura [ 278]
Gen. et sp. nov
Valid
Nel et al.
Permian (Kungurian)
Koshelevka Formation
Russia
A protozygopteran . The type species is A. permiana .
Balticoagrion [ 279]
Gen. et sp. nov
Valid
Bechly
Eocene
Europe
A damselfly found in Baltic amber . The type species is Balticoagrion paulyi .
Decoraeshna [ 280]
Gen. et sp. nov
Valid
Li et al.
Early Cretaceous
Yixian Formation
China
A progobiaeshnid dragonfly . The type species is Decoraeshna preciosus .
Engellestes [ 278]
Gen. et sp. nov
Valid
Nel et al.
Permian (Kungurian)
Koshelevka Formation
Russia
A protozygopteran . The type species is E. chekardensis .
Epilestes angustapterix [ 278]
Sp. nov
Valid
Nel et al.
Late Permian
Russia
A protozygopteran.
Germanostenolestes [ 281]
Gen. et sp. nov
Valid
Nel & Fleck
Middle Miocene
Germany
A sieblosiid epiproctophoran odonate . The type species is Germanostenolestes lutzi .
Karatawia sinica [ 282]
Sp. nov
Valid
Li, Nel & Ren in Li et al.
Middle Jurassic
Jiulongshan Formation
China
A campterophlebiid odonate ,a species of Karatawia .
Kennedya azari [ 278]
Sp. nov
Valid
Nel et al.
Permian (Kungurian)
Koshelevka Formation
Russia
A protozygopteran.
Kennedya ivensis [ 278]
Sp. nov
Valid
Nel et al.
Late Permian
Russia
A protozygopteran.
Kennedya pritykinae [ 278]
Sp. nov
Valid
Nel et al.
Late Permian
Russia
A protozygopteran.
Luiseia [ 278]
Gen. et sp. nov
Valid
Nel et al.
Carboniferous/Permian boundary
Bursum Formation
United States
A protozygopteran. The type species is L. breviata .
Mongoliaeshna exiguusens [ 280]
Sp. nov
Valid
Li et al.
Early Cretaceous
Yixian Formation
China
A progobiaeshnid dragonfly , a species of Mongoliaeshna .
Mongoliaeshna hadrens [ 280]
Sp. nov
Valid
Li et al.
Early Cretaceous
Yixian Formation
China
A progobiaeshnid dragonfly , a species of Mongoliaeshna .
Oligomazon [ 283]
Gen. et comb. nov
Valid
Zessin & Brauckmann
Carboniferous (Pennsylvanian )
United States
A member of Meganisoptera , a paralogid ; a new genus for "Oligotypus " makowskii Carpenter & Richardson (1971).
Paragilsonia [ 284]
Gen. et sp. nov (but see notes)
Valid
Zhang, Hong & Su in Su et al.
Carboniferous (Namurian )
China
A member of Odonatoptera ; the type species is Paragilsonia orientalis . Li et al. (2013) synonymized Paragilsonia with the genus Tupus Sellards (1906); however, they retained P. orientalis as a distinct species within the latter genus.[ 285]
Paralogobora [ 283]
Gen. et sp. nov
Valid
Zessin & Brauckmann
Early Permian
Czech Republic
A member of Odonata . The type species is Paralogobora guentherpetersi .
Permolestes sheimogorai [ 278]
Sp. nov
Valid
Nel et al.
Late Permian
Russia
A protozygopteran.
Permolestes soyanaiensis [ 278]
Sp. nov
Valid
Nel et al.
Late Permian
Russia
A protozygopteran.
Progoneura grimaldii [ 278]
Sp. nov
Valid
Nel et al.
Early Permian
Wellington Formation
United States
A protozygopteran.
Pseudocymatophlebia boda [ 286]
Sp. nov
Valid
Li, Nel & Ren in Li et al.
Early Cretaceous
China
An aktassiid odonate , a species of Pseudocymatophlebia .
Sinierasiptera [ 284]
Gen. et sp. nov (but see notes)
Valid
Zhang, Hong & Su in Su et al.
Carboniferous (Namurian )
China
A member of Odonatoptera ; the type species is Sinierasiptera jini . Li et al. (2013) synonymized Sinierasiptera with the genus Erasipterella Brauckmann (1983); however, they retained S. jini as a distinct species within the latter genus.[ 285]
Sinojagoria cancellosa [ 280]
Sp. nov
Valid
Li et al.
Early Cretaceous
Yixian Formation
China
A gomphaeschnid dragonfly , a species of Sinojagoria .
Sinojagoria magna [ 280]
Sp. nov
Valid
Li et al.
Early Cretaceous
Yixian Formation
China
A gomphaeschnid dragonfly , a species of Sinojagoria .
Solikamptilon pectinatus [ 278]
Sp. nov
Valid
Nel et al.
Late Permian
Russia
A protozygopteran.
Orthopterida
Raphidioptera
Other
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Anebos [ 297]
Gen. et sp. nov
Valid
Garwood et al.
Late Pennsylvanian
Assise de Montceau Formation
France
A polyneopteran insect of uncertain affinities. The type species is Anebos phrixos .
Anebos phrixos
Bechala [ 298]
Gen. et sp. nov
Valid
Ilger & Brauckmann
Late Carboniferous
Germany
A member of Odonatoptera , assigned to the new family Bechalidae . Originally interpreted as a megasecopteran .[ 299] Type species is Bechala sommeri .
Bruayaphis [ 300]
Gen. et sp. nov
Valid
Nel, Szwedo & Roques in Nel et al.
Late Carboniferous
France
A member of Paraneoptera of uncertain phylogenetic placement. The type species is Bruayaphis oudardi .
Cucullistriga [ 301]
Gen. et sp. nov
Valid
Aristov & Rasnitsyn
Kungurian
Russia
A member of Grylloblattida/Eoblattida belonging to the family Idelinellidae ; a relative of ice crawlers or cockroaches .[ 302] Type species is Cucullistriga cucullata .
Darekia sanguinea [ 303]
Gen. et sp. nov
Valid
Prokop et al.
Upper Carboniferous
Upper Silesian Coal Basin
Poland
A stem winged insect.
Eochauliodes [ 304]
Gen. et sp. nov
Valid
Liu et al.
Middle Jurassic (Bathonian-Callovian)
Jiulongshan Formation
China
A fishfly . The type species is Eochauliodes striolatus .
Eospilopsyllus [ 305]
Gen. et sp. nov
Valid
Beaucournu & Perrichot in Perrichot, Beaucournu & Velten
Early Miocene
Dominican Republic
A pulicid flea found in Dominican amber . The type species is Eospilopsyllus kobberti .
Geinitzia aristovi [ 306]
sp nov
Valid
Cui, Storozhenko & Ren
Middle Jurassic
Jiulongshan Formation
China
A geinitziid , a member of Polyneoptera /Gryllones belonging to the group Reculida ;[ 302] a species of Geinitzia .
Gymnopollisthrips [ 307]
Gen. et 2 sp. nov.
Valid
Peñalver, Nel & Nel in Peñalver et al.
Early Cretaceous
Spain
A melanthripid thrip . Genus contains two species: Gymnopollisthrips minor and G. maior .
Liadotaulius daohugouensis [ 308]
sp nov
Valid
Wu & Huang
Middle Jurassic
Daohugou Beds
China
A caddisfly , a species of Liadotaulius .
Netoxena [ 309]
Nom. nov
Valid
Sohn in Sohn et al.
Early Cretaceous (late Aptian)
Crato Formation
Brazil
A lepidopteran , possibly a member of Eolepidopterigidae ; a replacement name for Xena Martins-Neto (1999).
Pseudopulex [ 310]
Gen. et 2 sp. nov
Valid
Gao, Shih, & Ren in Gao et al.
Middle Jurassic to Early Cretaceous
China
A flea-like insect. The genus contains 2 species: P. jurassicus and P. magnus .
Rasstriga [ 301]
Gen. et sp. nov
Valid
Aristov & Rasnitsyn
Upper Carboniferous (Desmoinesian)
United States
A member of Grylloblattida/Eoblattida probably belonging to the family Idelinellidae ; a relative of ice crawlers or cockroaches .[ 302] Its type species is Rasstriga americana .
Scutistriga [ 301]
Gen. et sp. nov
Valid
Aristov & Rasnitsyn
Kungurian
Russia
A member of Grylloblattida/Eoblattida belonging to the family Idelinellidae ; a relative of ice crawlers or cockroaches .[ 302] Its type species is Scutistriga scutata .
Sharasialis [ 311]
Gen. et sp. nov
Valid
Ponomarenko
Late Jurassic
Mongolia
An alderfly . The type species is Sharasialis fusiformis .
Sinoprotodiplatys [ 312]
Gen. et sp. nov
Valid
Nel et al.
Early Cretaceous
Yixian Formation
China
An earwig belonging to the family Protodiplatyidae . The type species is Sinoprotodiplatys zhangi .
Strigulla [ 301]
Gen. et comb. nov
Valid
Aristov & Rasnitsyn
Kungurian
Russia
A member of Grylloblattida/Eoblattida belonging to the family Idelinellidae ; a relative of ice crawlers or cockroaches .[ 302] A new genus for "Euryptilon " cuculiophoris (Aristov, 2002).
Strudiella [ 313]
Gen. et sp. nov.
Valid
Garrouste et al.
Late Devonian
Belgium
The first complete Late Devonian insect, probably terrestrial. The type species is Strudiella devonica .
Zdenekia silesiensis [ 303]
Species
Valid
Prokop et al.
Upper Carboniferous
Upper Silesian Coal Basin
Poland
A stem winged insect.
General research
Trilobites
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Abanomocarella [ 318]
Gen. et sp. et comb. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Ptychopariida belonging to the family Anomocarellidae . Genus includes new species A. quadrata , as well as A. humilis (Resser & Endo in Endo & Resser, 1937) and A. pyriformis (Resser & Endo in Endo & Resser, 1937).
Acmarhachis kindlei [ 319]
Sp nov
Valid
Westrop & Eoff
Late Cambrian
Cow Head Group
Canada
An agnostid , a species of Acmarhachis .
Acmarhachis whittingtoni [ 319]
Sp nov
Valid
Westrop & Eoff
Late Cambrian
Cow Head Group
Canada
An agnostid , a species of Acmarhachis .
Angsiduoa jiulongshanensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Ptychopariida belonging to the family Ignotogregatidae .
Angsiduoa shanxiensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Ptychopariida belonging to the family Ignotogregatidae .
Angsiduoa transversa [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Ptychopariida belonging to the family Ignotogregatidae .
Anomocarella? antiqua [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Ptychopariida belonging to the family Anomocarellidae .
Anomocarella jiulongshanensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Ptychopariida belonging to the family Anomocarellidae .
Anomocarella similis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Ptychopariida belonging to the family Anomocarellidae .
Aponileus aasei [ 320]
Sp nov
Valid
Adrain & McAdams
Early Ordovician (late Floian)
Fillmore Formation
United States
A bathyurid trilobite, a species of Aponileus .
Aponileus belkaae [ 320]
Sp nov
Valid
Adrain & McAdams
Early Ordovician (late Floian)
Wah Wah Formation
United States
A bathyurid trilobite, a species of Aponileus .
Aponileus laikaae [ 320]
Sp nov
Valid
Adrain & McAdams
Early Ordovician (late Floian)
Fillmore Formation
United States
A bathyurid trilobite, a species of Aponileus .
Aponileus strelkaae [ 320]
Sp nov
Valid
Adrain & McAdams
Early Ordovician (late Floian)
Wah Wah Formation
United States
A bathyurid trilobite, a species of Aponileus .
Aponileus ugolekae [ 320]
Sp nov
Valid
Adrain & McAdams
Early Ordovician (late Floian)
Wah Wah Formation
United States
A bathyurid trilobite, a species of Aponileus .
Aponileus? veterokae [ 320]
Sp nov
Valid
Adrain & McAdams
Early Ordovician (late Floian)
Wah Wah Formation
United States
A bathyurid trilobite, a possible species of Aponileus .
Archaeagnostus typicus [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Peronopsidae .
Archegonus (Phillibole) nehdenensis africanus [ 321]
Subsp. nov
Valid
Hahn, Müller & Becker
Carboniferous (Mississippian )
Morocco
A member of Phillipsiidae belonging to the subfamily Archegoninae , a subspecies of Archegonus nehdenensis .
Asaphellus cuervoae [ 322]
Sp. nov
Disputed
Corbacho & López-Soriano
Ordovician (Floian )
Upper Fezouata Formation
Morocco
A member of Asaphidae , a species of Asaphellus . Gutiérrez-Marco et al. (2017) considered this species to be likely based on poorly preserved and probably retouched specimens of Asaphellus stubbsi .[ 323]
Astycoryphe arduinnae [ 324]
Sp nov
Valid
Van Viersen, van Rossum & Prescher
Middle Devonian
Belgium
A member of Proetida .
Australoscutellum iota [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Australoscutellum .
Australoscutellum microps [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Australoscutellum .
Australoscutellum pulex [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Australoscutellum .
Australoscutellum trica [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Australoscutellum .
Baltagnostus yaojiayuensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Diplagnostidae .
Baojingia (Eobaojingia) [ 318]
Subgen. et 2 sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Lisaniidae . The subgenus includes Baojingia (Eobaojingia) dubia and Baojingia (Eobaojingia) ezhuangensis , as well as Baojingia miaogouensis Mong in Zhou et al. (1977).
Borenoria [ 325]
gen et sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite. The type species is Borenoria cirrita .
Bumastoides graffhami [ 326]
Sp nov
Valid
Carlucci et al.
Ordovician
United States
An illaenid trilobite, a species of Bumastoides .
Bumastoides kimmswickensis [ 326]
Sp nov
Valid
Carlucci et al.
Ordovician
United States
An illaenid trilobite, a species of Bumastoides .
Bumastoides moundensis [ 326]
Sp nov
Valid
Carlucci et al.
Ordovician
United States
An illaenid trilobite, a species of Bumastoides .
Calymene harringtoni [ 327]
Sp. nov
Valid
Tortello in Tortello et al.
Early Silurian
Vargas Peña Formation
Paraguay
A species of Calymene .
Chlupacula (Chlupacula) erfoudensis [ 321]
Sp. nov
Valid
Hahn, Müller & Becker
Carboniferous (Mississippian )
Morocco
A member of Phillipsiidae belonging to the subfamily Archegoninae , a species of Chlupacula .
Crepicephalina exsculpta [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Crepicephalina obscura [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Crepicephalina pulchra [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Crepicephalina? rara [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Crepicephalina woqishanensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Cummingella boikoi [ 328]
sp nov
Valid
Mychko
Sakmarian
Russia
A phillipsiid , a species of Cummingella .
Cyrtoproetus (Crassibole) acrops [ 321]
Sp. nov
Valid
Hahn, Müller & Becker
Carboniferous (Mississippian )
Morocco
A member of Phillipsiidae belonging to the subfamily Archegoninae , a species of Cyrtoproetus .
Dalejeproetus owensi [ 329]
Sp nov
Valid
Johnson & Fortey
Early Devonian (Pragian)
Ihandar Formation
Morocco
A proetid trilobite, a species of Dalejeproetus .
Dalejeproetus sagaouii [ 329]
Sp nov
Valid
Johnson & Fortey
Early Devonian (Pragian)
Ihandar Formation
Morocco
A proetid trilobite, a species of Dalejeproetus .
Dameselloides [ 318]
Gen. et sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Damesellidae . Genus includes new species D. sinensis .
Danjiangella shandongensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Dechenelloides tazoultensis [ 321]
Sp. nov
Valid
Hahn, Müller & Becker
Carboniferous (Mississippian )
Morocco
A member of Phillipsiidae belonging to the subfamily Archegoninae , a species of Dechenelloides .
Decoroscutellum planes [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Decoroscutellum .
Derikaspis conicus [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Derikaspis ruichengensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Diacoryphe (Archaeocoryphe) maiderensis [ 321]
Sp. nov
Valid
Hahn, Müller & Becker
Carboniferous (Mississippian )
Morocco
A member of Phillipsiidae belonging to the subfamily Archegoninae , a species of Diacoryphe .
Dindymene whittingtoni [ 330]
Sp nov
Valid
Owen & Bruton
Late Ordovician
United States
A species of Dindymene .
Dorypyge cylindrica [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Dorypyge zhangxianensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Dunopyge seiberti [ 331]
Sp. nov
Valid
Basse & Müller
Early Devonian
Seifen Beds
Germany
A member of Acastidae belonging to the subfamily Asteropyginae , a species of Dunopyge .
Eilura trapezialis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Elaphraella? taebaeksanensis [ 332]
Sp. nov
Valid
Park & Choi
Cambrian (middle Furongian )
Sesong Formation
South Korea
A member of the family Shumardiidae .
Emmrichella punctata [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Ptychopariidae .
Eokoldinia [ 318]
Gen. et sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Plethopeltidae . Genus includes new species E. porifera .
Eomapania [ 318]
Gen. et sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Mapaniidae . Genus includes new species E. subquadrata .
Eoscutellum annosum [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Eoscutellum .
Eymekops pristina [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Eymekops transversa [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Grandioculus macroculus [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Grandioculus (Protohedina) quadraticephala [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Gumunsoia [ 333]
Gen. et sp. nov
Valid
Park, Sohn & Choi
Cambrian (middle Furongian )
Sesong Formation
South Korea
A member of Leiostegioidea belonging to the family Kaolishaniidae . Genus includes new species G. triangularis .
Gushanella [ 318]
Gen. et 2 sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Solenopleuridae . Genus includes new species G. quadrata and G. truncata .
Heyelingella crassa [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Honania porifera [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Huayuanaspis angustilimbata [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Idioura brevispina [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Illaenoscutellum tryo [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Illaenoscutellum .
Inouyella rara [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Inouyella shiliuyuanensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Inouyia (Bulbinouyia) [ 318]
Subgen. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Inouyiidae . The subgenus includes Inouyia kangzhuangensis (Zhang & Wang, 1985) and I. lubrica (Zhou in Zhou et al. , 1982).
Inouyia latilimbata [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Iranoleesia pustulosa [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Iranoleesia (Proasaphiscina) rogosa [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Japonoscutellum borenorense [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Japonoscutellum .
Japonoscutellum crassum [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Japonoscutellum .
Japonoscutellum diascetum [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Japonoscutellum .
Japonoscutellum gephyricum [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Japonoscutellum .
Japonoscutellum guttulatum [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Japonoscutellum .
Japonoscutellum serangicum [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Japonoscutellum .
Japonoscutellum strabo [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Japonoscutellum .
Jiulongshania shandongensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Diceratocephalidae .
Koptura (Eokoptura) [ 318]
Subgen. et sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Anomocaridae . The subgenus includes new species Koptura (Eokoptura) bella , as well as Koptura quadrata (Resser & Endo in Endo & Resser, 1937).
Koptura intermedia [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Anomocaridae .
Kosovopeltis trepida [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Kosovopeltis .
Kosovopeltis xynon [ 325]
sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite, a species of Kosovopeltis .
Lepidoproetus lahceni [ 329]
Sp nov
Valid
Johnson & Fortey
Early Devonian (Pragian)
Ihandar Formation
Morocco
A proetid trilobite, a species of Lepidoproetus .
Lepidoproetus maharchianus [ 329]
Sp nov
Valid
Johnson & Fortey
Early Devonian (Pragian)
Ihandar Formation
Morocco
A proetid trilobite, a species of Lepidoproetus .
Lepidoproetus splendens [ 329]
Sp nov
Valid
Johnson & Fortey
Early Devonian (Pragian)
Ihandar Formation
Morocco
A proetid trilobite, a species of Lepidoproetus .
Liowutaishania laevigata [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Lisania elliptica [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Longiproetus meiopsifius [ 324]
Sp nov
Valid
Van Viersen, van Rossum & Prescher
Middle Devonian
Belgium
A member of Proetida .
Lunania [ 318]
Gen. nov
Junior homonym
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Menocephalidae . The generic name is preoccupied by Lunania Chow (1957).
Maotunia transversa [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Mapania (Mapanella) [ 318]
Subgen. et 2 sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Mapaniidae . The subgenus includes new species M. (Mapanella) transversa and M. (Mapanella) prisca , as well as Mapania blackwelderi (Resser & Endo in Endo & Resser, 1937).
Mapania subcylindrica [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Mapanopsis conicus [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Maurotarion megacephalum [ 334]
Sp. nov.
Valid
Rustán & Vaccari
Early Devonian (Pragian )
Talacasto Formation
Argentina
An otarionine aulacopleurid , a species of Maurotarion .
Megagraulos similis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Menocephalina [ 318]
Gen. et sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Menocephalidae . Genus includes new species M. longispinosa .
Menocephalites (Hutoushania) [ 318]
Subgen. et 2 sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Menocephalidae . The subgenus includes Menocephalites (Hutoushania) triangulata and M. (Hutoushania) zaozhuangansis .
Menocephalites porosus [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Menocephalites primitivus [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Menocephalites simplex [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Metanomocarella primitiva [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Micagnostus semicircularis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Mirrabooka [ 325]
gen et sp nov
Valid
Holloway and Lane
Silurian
Australia
A scutelluid trilobite. The type species is Mirrabooka harryi .
Monkaspis conica [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Ohleum magreani [ 335]
Sp nov
Valid
Taghon, Bonino & Mottequin
Devonian (Eifelian )
Jemelle Formation
Belgium
A member of Lichida belonging to the family Lichidae .
Pagetides qianensis [ 336]
Sp nov
Valid
Zhang & Clarkson
Cambrian
China
An eodiscoid trilobite, a species of Pagetides .
Paleonelsonia granulosa [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Ptychopariidae .
Paleonelsonia sulcata [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Ptychopariidae .
Paracoosia langgongsiensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Anomocaridae .
Paralevisia subglobosa [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Parayujinia pulchra [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Peishania abnormis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Peishania obscura [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Peronopsis jiulongshanensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Peronopsis universalis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Persia [ 337]
Gen. et sp. nov
Junior homonym
Lerosey-Aubril
Carboniferous (Tournaisian)
Iran
The type species is Persia praecox . The generic name is preoccupied by Persia Repin (1996); Lerosey-Aubril & Deshmukh (2022) coined a replacement name Persiax .[ 338]
Pilletina seifenensis [ 331]
Sp. nov
Valid
Basse & Müller
Early Devonian
Seifen Beds
Germany
A member of Acastidae belonging to the subfamily Asteropyginae , a species of Pilletina .
Pingluia granulosa [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Plectocrania elegans [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Plectocrania intermedia [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Plesigangderria latilimbata [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Podoliproetus mirdani [ 329]
Sp nov
Valid
Johnson & Fortey
Early Devonian (Pragian)
Ihandar Formation
Morocco
A proetid trilobite, a species of Podoliproetus .
Podoliproetus sinespina [ 329]
Sp nov
Valid
Johnson & Fortey
Early Devonian (Pragian)
Ihandar Formation
Morocco
A proetid trilobite, a species of Podoliproetus .
Praeymekops [ 318]
Gen. et sp. et comb. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Proasaphiscidae . Genus includes new species P. minutus , as well as P. sinensis (Mansuy, 1916).
Proasaphiscus laiwuensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Proetina ihamadii [ 329]
Sp nov
Valid
Johnson & Fortey
Early Devonian (Pragian)
Ihandar Formation
Morocco
A proetid trilobite, a species of Proetina .
Pseudoolenoides fossilmountainensis [ 339]
Sp nov
Valid
Adrain, McAdams & Karim
Middle Ordovician
Kanosh Formation
United States
A bathyurid , a species of Pseudoolenoides .
Pseudoolenoides oilcreekensis [ 339]
Sp nov
Valid
Adrain, McAdams & Karim
Middle Ordovician
Oil Creek Formation
United States
A bathyurid , a species of Pseudoolenoides .
Pseudoolenoides pogonipensis [ 339]
Sp nov
Valid
Adrain, McAdams & Karim
Middle Ordovician
Kanosh Formation
United States
A bathyurid , a species of Pseudoolenoides .
Pseudopeishania [ 318]
Gen. et 3 sp. et comb. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Ptychopariida belonging to the family Anomocarellidae . Genus includes new species P. brevis , P. similis and P. woqishanensis , as well as P. angustata (Endo, 1944).
Pseudoperonopsis elegans [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Pseudophillipsia parvizii [ 337]
Sp. nov
Valid
Lerosey-Aubril
Permian
Iran
A species of Pseudophillipsia sensu lato .
Pseudotaitzuia angustilimbata [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Asaphida belonging to the family Ordosiidae.
Pseudotaitzuia rara [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Asaphida belonging to the family Ordosiidae.
Pudoproetus zhorae [ 321]
Sp. nov
Valid
Hahn, Müller & Becker
Carboniferous (Mississippian )
Morocco
A member of Proetidae , a species of Pudoproetus .
Raragnostus intermedius [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Raymondaspis grandigena [ 340]
sp nov
Valid
Stein & Bergström
Middle Ordovician
Sweden
A styginid , a species of Raymondaspis .
Reediella mitchelli tafilaltensis [ 321]
Subsp. nov
Valid
Hahn, Müller & Becker
Carboniferous (Mississippian )
Morocco
A member of Phillipsiidae belonging to the subfamily Bollandiinae , a subspecies of Reediella mitchelli .
Ruichengocephalus [ 318]
Gen. et sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Ptychopariidae . Genus includes new species R. typicus .
Saimachia yichangensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Schopfaspis? graciai [ 341]
sp nov
Valid
Esteve et al. .
Middle Cambrian
Spain
An alokistocarid , possibly a species of Schopfaspis .
Sdzuyella conica [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Ptychopariida belongig to the family Holocephalidae .
Securifrons [ 325]
gen et comb nov
Valid
Holloway and Lane
Ordovician to Silurian
Scutelluid trilobite. species- S. borealis , S. cunctatum , S. major and ?S. umbonatus .
Shiliuyuania [ 318]
Gen. et 2 sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Crepicephalidae . Genus includes new species S. cylindrica and S. quadrata .
Solenoparia zhangxianensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Solenoparina bella [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Sphaerophthalmus arcus [ 342]
Sp nov
Valid
Høyberget & Bruton
Cambrian (Furongian )
Alum Shale Formation
Norway
A member of the family Olenidae .
Sudanomocarina elliptica [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Sudanomocarina hutoushanensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Symphysops stevaninae [ 343]
Sp. nov
Valid
López-Soriano & Corbacho
Late Ordovician
Lower Ktaoua Formation
Morocco
A member of Cyclopygidae , a species of Symphysops .
Szeaspis (Pseudoszeaspis) [ 318]
Subgen. et sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Ptychopariida belonging to the family Anomocarellidae . The subgenus includes new species Szeaspis (Pseudoszeaspis) conicus , as well as Szeaspis huananensis Peng, Babcock & Lin (2004) and Szeaspis pustulosa (Resser & Endo in Endo & Resser, 1937).
Szeaspis crassus [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Ptychopariida belonging to the family Anomocarellidae .
Taitzehoia huolianzhaiensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Damesellidae .
Teinistion jiulongshanense [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Damesellidae .
Teinistion quadratum [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Damesellidae .
Teinistion yangjiazhuangense [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Damesellidae .
Tomagnostus (Paratomagnostus) [ 318]
Subgen. et sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Ptychagnostidae . The subgenus includes Tomagnostus (Paratomagnostus) ezhuangensis .
Trachoparia elongata [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Trachoparia intermedia [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Trachoparia spinosa [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Triangulopyge [ 342]
Gen. et comb. nov
Valid
Høyberget & Bruton
Cambrian
Norway
Sweden
United Kingdom
A member of the family Olenidae . The type species is "Olenus" humilis Phillips (1848); genus also includes "Sphaerophthalmus " majusculus Linnarsson (1880) and T. major (Lake, 1913).
Tsunyidiscus pengshuiensis [ 336]
Sp nov
Valid
Zhang & Clarkson
Cambrian
China
An eodiscoid trilobite, a species of Tsunyidiscus .
Ulania concava [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Utagnostus primitivus [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Clavagnostidae .
Utagnostus sulcatus [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Clavagnostidae .
Woqishanaspis [ 318]
Gen. et comb. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Ignotogregatidae . Genus includes W. trapezoidalis (Lin, 1984).
Wutaishanaspis yangjiazhuangensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Xenolorenzella granulosa [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Yichengocephalus [ 318]
Gen. et sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Solenopleuridae . Genus includes new species Y. latus .
Yongwolia quadrata [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Yushugouia ezhuangensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Zhangxiaspis [ 318]
Gen. et sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Ptychopariidae . Genus includes new species Z. subcylindrica .
Zhongtiaoshanella [ 318]
Gen. et sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Crepicephalidae . Genus includes new species Z. brevispinosa .
Zhongweia convexa [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
Zhujia yangjiazhuangensis [ 318]
Sp. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of Ptychopariida belonging to the family Anomocarellidae .
Zichuanaspis [ 318]
Gen. et 2 sp. et comb. nov
Valid
Yuan et al.
Cambrian
Changhia Formation
China
A member of the family Leiostegiidae . Genus includes Zichuanaspis acerius (Walcott, 1905) and Z. catinus (Resser & Endo in Endo & Resser, 1937), as well as new species Z. yangjiazhuangensis and possibly Z? latilimbata .
Others
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Australimicola [ 344]
Gen. et sp.
Valid
Paterson, García-Bellido and Edgecombe
Early Cambrian
The Emu Bay Shale Konservat-Lagerstätte
Australia
An artiopodan arthropod. The type species is Australimicola spriggi .
Kangacaris shui [ 345]
Sp. nov
Valid
Zhang, Fu & Dai
Lower Cambrian
Chengjiang lagerstätte
China
A species of Kangacaris .
Luohuilinella [ 346]
Gen. et sp. nov
Valid
Zhang, Fu & Dai
Lower Cambrian
Chengjiang Lagerstätte
China
A possible member of Xandarellida . The type species is Luohuilinella rarus .
Nereocaris [ 347]
Gen. et sp. nov
Valid
Legg et al.
Cambrian
Burgess Shale Formation
Canada
A bivalved arthropod, one of the basalmost known arthropods. The type species is Nereocaris exilis .
Pleuralata [ 348]
Gen. et sp. nov
Valid
McCoy, Strother and Briggs
Silurian (Llandovery )
Tuscarora Formation
United States
An arthropod of uncertain phylogenetic affinities, a probable tracemaker of Arthrophycus alleghaniensis trace fossils. The type species is Pleuralata spinosa .
Scolopocryptops simojovelensis [ 349]
Sp. nov
Valid
Edgecombe et al.
Miocene
Mexico
A scolopocryptopid scolopendromorph centipede , a species of Scolopocryptops .
Sinoleperditia hamadai [ 350]
Sp. nov
Valid
Tanaka, Ono and Maeda
Early Devonian
Japan
A member of Leperditicopida (a group of arthropods of uncertain affinities, classified as ostracods or excluded from this group by different authors), a species of Sinoleperditia .
Sinoleperditia huyeni [ 351]
Sp. nov
Valid
Tanaka, Phong and Komatsu
Early Devonian
Vietnam
A member of Leperditicopida , a species of Sinoleperditia .
Squamacula buckorum [ 344]
Species
Valid
Paterson, García-Bellido and Edgecombe
Early Cambrian
The Emu Bay Shale Konservat-Lagerstätte
Australia
An artiopodan arthropod, a species of Squamacula .
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^ a b Jörg Wunderlich; Patrick Müller (2018). "Reflections on the origin of the orb web, on the evolution and the relationships of selected spiders (Araneae): the RTA-clade, the superfamily Leptonetoidea as well as on the orb-weaving spiders (Araneoidea and Deinopoidea), based on "missing links" in Cretaceous Burmese amber; descriptions of new taxa, and notes on the spider fauna in Burmite" (PDF) . In Jörg Wunderlich (ed.). Beiträge zur Araneologie, 11. Fossil spiders (Araneae) in Cretaceous Burmese amber . Publishing House Joerg Wunderlich. pp. 1–177. ISBN 978-3-931473-16-7 .
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^ a b c d Carrie E. Schweitzer; Rodney M. Feldmann; Silvio Casadío; Martin Rodriguez Raising (2012). "Eocene Decapod Crustacea (Thalassinidea and Brachyura) from Patagonia, Argentina". Annals of Carnegie Museum . 80 (3): 173–186. doi :10.2992/007.080.0301 . S2CID 86083986 .
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^ Jun-Jie Gu; Xin Yang; Rong Huang; Guijun Yang; Yanli Yue; Dong Ren (2021). "New species and material of Hagloidea (Insecta, Ensifera) from the Yanliao biota of China" . ZooKeys (1033): 183–190. Bibcode :2021ZooK.1033..183G . doi :10.3897/zookeys.1033.63571 . PMC 8084857 . PMID 33958925 .
^ Olivier Béthoux; Junjie Gu; Yanli Yue; Dong Ren (2012). "Miamia maimai n. sp., a new Pennsylvanian stem-orthopteran insect, and a case study on the application of cladotypic nomenclature" . Fossil Record . 15 (2): 103–113. doi :10.1002/mmng.201200008 .
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^ You-chong Hong; Zhen-yu Li (2012). "The discovery of a new family Sunopteridae fam. nov. (Insecta: Protorthoptera) from Shaanxi Province, China" . Geological Bulletin of China . 31 (5): 647–652.
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^ Xingyue Liu; Yongjie Wang; Chungkun Shih; Dong Ren; Ding Yang (2012). "Early Evolution and Historical Biogeography of Fishflies (Megaloptera: Chauliodinae): Implications from a Phylogeny Combining Fossil and Extant Taxa" . PLOS ONE . 7 (7): e40345. Bibcode :2012PLoSO...740345L . doi :10.1371/journal.pone.0040345 . PMC 3391272 . PMID 22792287 .
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^ Yingying Cui; Sergey Yu Storozhenko; Dong Ren (2012). "New and little-known species of Geinitziidae (Insecta: Grylloblattida) from the Middle Jurassic of China, with notes on taxonomy, habitus and habitat of these insects". Alcheringa: An Australasian Journal of Palaeontology . 36 (2): 251–261. Bibcode :2012Alch...36..251C . doi :10.1080/03115518.2012.628806 . S2CID 129794581 .
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