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角蟾属
	山角蟾（Megophrys montana）
科学分类
界：	动物界 Animalia
门：	脊索动物门 Chordata
纲：	两栖纲 Amphibia
目：	无尾目 Anura
科：	角蟾科 Megophryidae
属：	角蟾属 Megophrys; Kuhl & van Hasselt, 1822
模式種
	山角蟾; Megophrys montana; Kuhl & van Hasselt，1822
異名
	Atympanophrys Tian & Hu, 1983 ; Borealophrys Fei & Ye, 2016 ; 婆罗蟾属 Borneophrys Delorme, Dubois, Grosjean & Ohler, 2006 ; Ceratophryne Schlegel, 1858 ; Gigantophrys Fei & Ye, 2016 ; Liuophrys Fei & Ye, 2016 ; Megalophrys Kuhl & Van Hasselt, 1822 ; Megalophys Gray, 1842 ; Mogophrys Kuhl & Van Hasselt, 1822 ; 拟角蟾属 Ophryophryne Boulenger, 1903 ; Pelobatrachus Beddard, 1908 ;

角蟾屬（学名：Megophrys）是无尾目角蟾科下的一个属。分布于东南亚地区。本属原包含多个物种，现大部分种类已被归入布角蟾属（Boulenophrys）。

下属物种

截止2023年3月，根据 Amphibian Species of the World 中的记录，本属包含以下5种：^[1]

亚齐角蟾 Megophrys acehensis Munir, Nishikawa, Hamidy, and Smith, 2021
Megophrys ancrae Mahony, Teeling & Biju, 2013^[2]
Megophrys (Xenophrys) awuh ^[2]
Megophrys caobangensis Nguyen, Pham, Nguyen, Luong & Ziegler, 2020^[3]
陈氏角蟾 Megophrys cheni (Wang & Yang, 2014)，基异名：Xenophrys cheni Wang & Yang, 2014^[4]
Megophrys (Xenophrys) dzukou ^[2]
Megophrys (Panophrys) fansipanensis Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Lương & Rowley, 2018^[5]
Megophrys frigida Tapley, Cutajar, Nguyen, Portway, Mahony, Nguyen, Harding, Luong & Rowley, 2020^[6]
喜马拉雅角蟾 Megophrys himalayana Mahony, Kamei, Teeling & Biju, 2018^[7]
井冈山角蟾 Megophrys jinggangensis (Wang, 2012)，基异名：Xenophrys jinggangensis Wang, 2012^[8]
加里曼丹角蟾 Megophrys kalimantanensis Munir, Hamidy, Matsui, Iskandar, Sidik & Shimada, 2019^[9]
Megophrys lancip Munir, Hamidy, Farajallah, and Smith, 2018^[10]
Megophrys lini (Wang & Yang, 2014)，基异名：Xenophrys lini Wang & Yang, 2014^[4]
泸水角蟾 Megophrys lushuiensis Shi, Li, Zhu, Jiang, Jiang & Wang, 2021^[11]
山角蟾 Megophrys montana Kuhl and Van Hasselt, 1822
Megophrys (Xenophrys) numhbumaeng Mahony, Kamei, Teeling & Biju, 2020^[12]
雨神角蟾 Megophrys ombrophila Messenger & Dahn^[13]
Megophrys oropedion Mahony, Teeling & Biju, 2013^[2]
Megophrys parallela Inger and Iskandar, 2005
红股异角蟾 Megophrys (Xenophrys) rubrimera Tapley, Cutajar, Mahony, Nguyen, Dau, Nguyen, Luong & Rowley, 2017^[14]
南苏门答腊角蟾 Megophrys selatanensis Munir, Nishikawa, Hamidy, and Smith, 2021^[15]
Megophrys (Xenophrys) serchhipii (Mathew & Sen, 2007) ^[2]
沙坪角蟾 Megophrys shapingensis Liu, 1950
舜皇角蟾 Megophrys shunhuangensis Wang, Deng, Liu, Wu & Liu, 2019，分布于湖南、广西。
Megophrys vegrandis Mahony, Teeling & Biju, 2013^[2]
Megophrys (Xenophrys) zunhebotoensis (Mathew & Sen, 2007)^[2]

分类

角蟾属分类存在争议。一种意见认为无耳蟾属(Atympanophrys)、短腿蟾属(Brachytarsophrys)、拟角蟾属(Ophryophryne)、泛角蟾属(Panophrys)及异角蟾属(Xenophrys)都应视为角蟾属的同物异名^[16]。

2017年以后的多项研究都对角蟾属提出了不同的分类方案，一种是分为短腿蟾属、拟角蟾属、无耳蟾属、异角蟾属，同时将角蟾属限定于巽他古陆，婆罗蟾属则作为角蟾属的异名^[17]；另一种意见则将拟角蟾属、无耳蟾属、短腿蟾属、泛角蟾属和异角蟾属均降为亚属、并视为角蟾属的同物异名^[18]。

参考文献

^ Frost, Darrel R. 2023. Amphibian Species of the World: an online reference. Version 6.1 (21 March 2023). Electronic Database accessible at https://amphibiansoftheworld.amnh.org/Amphibia/Anura/Megophryidae/Megophryinae/Megophrys （页面存档备份，存于互联网档案馆）. American Museum of Natural History, New York, USA.
^ ^2.0 ^2.1 ^2.2 ^2.3 ^2.4 ^2.5 ^2.6 Mahony, Stephen, Emma C. Teeling & S. D. Biju. 2013. Three New Species of Horned Frogs, Megophrys (Amphibia: Megophryidae), from Northeast India, with a resolution to the identity of Megophrys boettgeri populations reported from the region. Zootaxa. 3722(2): 143-169.
^ Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Mai Luong and Thomas Ziegler. 2020. A New Species of Megophrys (Amphibia: Anura: Megophryidae) from Vietnam. Zootaxa. 4722(5); 401–422. DOI: 10.11646/zootaxa.4722.5.1
^ ^4.0 ^4.1 Yingyong Wang, Jian Zhao, Jianhuan Yang, Zhixin Zhou, Guoling Chen and Yang Liu. 2014. Morphology, Molecular Genetics, and Bioacoustics Support Two New Sympatric Xenophrys Toads (Amphibia: Anura: Megophryidae) in Southeast China. PLoS ONE. 9(4): e93075. DOI: 10.1371/journal.pone.0093075
^ Benjamin Tapley, Timothy Cutajar, Stephen Mahony, Chung T. Nguyen, Vinh Q. Dau, Anh M. Luong, Dzung T. Le, Tao T. Nguyen, Truong Q. Nguyen, Christopher Portway, Hào V. Lương and Jodi J. L. Rowley. 2018. Two New and Potentially Highly threatened Megophrys Horned Frogs (Amphibia: Megophryidae) from Indochina’s Highest Mountains. Zootaxa. 4508(3); 301–333. DOI: 10.11646/zootaxa.4508.3.1
^ Benjamin Tapley, Timothy Cutajar, Luan Thanh Nguyen, Christopher Portway, Stephen Mahony, Chung Thanh Nguyen, Luke Harding, Hao Van Luong and Jodi J. L. Rowley. 2020. A New Potentially Endangered Species of Megophrys (Amphibia: Megophryidae) from Mount Ky Quan San, north-west Vietnam. Journal of Natural History. 54(39-40); 2543-2575. DOI: 10.1080/00222933.2020.1856952
^ Stephen Mahony, Rachunliu G. Kamei, Emma C. Teeling and S. D. Biju. 2018. Cryptic Diversity within the Megophrys major Species Group (Amphibia: Megophryidae) of the Asian Horned Frogs: Phylogenetic Perspectives and A Taxonomic Revision of South Asian Taxa, with Descriptions of Four New Species. Zootaxa. 4523(1); 1–96. DOI: 10.11646/zootaxa.4523.1.1
^ Wang, Ying-yong, Tian-du Zhang, Jian Zhao, Yik-Hei Sung, Jian-huan Yang, Hong Pang & Zhong Zhang. 2012. Description of A New Species of the Genus Xenophrys Günther, 1864 (Amphibia: Anura: Megophryidae) from Mount Jinggang, China, based on Molecular and Morphological data. Zootaxa. (3546): 53-67.
^ Misbahul Munir, Amir Hamidy, Masafumi Matsui, Djoko T. Iskandar, Irvan Sidik and Tomohiko Shimada. 2019. A New Species of Megophrys Kuhl & Van Hasselt (Amphibia: Megophryidae) from Borneo allied to M. nasuta (Schlegel, 1858). Zootaxa. 4679(1); 1–24. DOI: 10.11646/zootaxa.4679.1.1
^ Misbahul Munir, Amir Hamidy, Achmad Farajallah and Eric N. Smith. 2018. A New Megophrys Kuhl and Van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa. 4442(3); 389–412. DOI: 10.11646/zootaxa.4442.3.3
^ 云南高黎贡山发现两栖动物一新种――泸水角蟾. 发现. [2023-07-12]. （原始内容存档于2023-07-12）.
^ Stephen Mahony, Rachunliu G. Kamei, Emma C. Teeling and S. D. Biju. 2020. Taxonomic Review of the Asian Horned Frogs (Amphibia: Megophrys Kuhl & Van Hasselt) of Northeast India and Bangladesh previously misidentified as M. parva (Boulenger), with Descriptions of Three New Species. Journal of Natural History. 54(1-4 - Amphibian taxonomy: Early 21st century case studies); DOI: 10.1080/00222933.2020.1736679
^ Kevin R. Messenger, Hollis A. Dahn, Yuanrui Liang, Peng Xie, Yong Wang and Changhu Lu. 2019. A New Species of the Genus Megophrys Gunther, 1864 (Amphibia: Anura: Megophryidae) from Mount Wuyi, China. Zootaxa. 4554(2); 561–583. DOI: 10.11646/zootaxa.4554.2.9
^ Benjamin Tapley, Timothy Cutajar, Stephen Mahony, Chung Thanh Nguyen, Vinh Quang Dau, Tao Thien Nguyen, Hao Van Luong and Jodi J. L. Rowley. 2017. The Vietnamese Population of Megophrys kuatunensis (Amphibia: Megophryidae) Represents A New Species of Asian Horned Frog from Vietnam and southern China. Zootaxa. 4344(3); 465–492. DOI: 10.11646/zootaxa.4344.3.3
^ Misbahul Munir, Kanto Nishikawa, Amir Hamidy and Eric N. Smith. 2021. Two New Species of Megophrys Kuhl and Van Hasselt (Amphibia: Megophryidae) from Sumatra, Indonesia. Zootaxa. 5057(4); 503-529. DOI: 10.11646/zootaxa.5057.4.3
^ Kai, Wang; Jinlong, Ren; Hongman, Chen; Zhitong, Lyu; Xianguang, Guo; Ke, Jiang; Jinmin, Chen; Jiatang, Li; Peng, Guo; Yingyong, Wang; Jing, Che. The updated checklists of amphibians and reptiles of China. Biodiversity Science. 2020-02-20, 28 (2): 189. doi:10.17520/biods.2019238.
^ Chen, Jin-Min; Poyarkov, Nikolay A.; Suwannapoom, Chatmongkon; Lathrop, Amy; Wu, Yun-He; Zhou, Wei-Wei; Yuan, Zhi-Yong; Jin, Jie-Qiong; Chen, Hong-Man; Liu, He-Qun; Nguyen, Truong Quang; Nguyen, Sang Ngoc; Duong, Tang Van; Eto, Koshiro; Nishikawa, Kanto; Matsui, Masafumi; Orlov, Nikolai L.; Stuart, Bryan L.; Brown, Rafe M.; Rowley, Jodi J.L.; Murphy, Robert W.; Wang, Ying-Yong; Che, Jing. Large-scale phylogenetic analyses provide insights into unrecognized diversity and historical biogeography of Asian leaf-litter frogs, genus Leptolalax (Anura: Megophryidae). Molecular Phylogenetics and Evolution. 2018-07, 124: 162–171. doi:10.1016/j.ympev.2018.02.020.
^ Mahony, Stephen; Foley, Nicole M.; Biju, S.D.; Teeling, Emma C. Evolutionary History of the Asian Horned Frogs (Megophryinae): Integrative Approaches to Timetree Dating in the Absence of a Fossil Record. Molecular Biology and Evolution. 2017-01-18: msw267. doi:10.1093/molbev/msw267.

外部链接

http://liangqi.sppchina.com/sppdb/1/82/^{[永久失效連結]}
http://www.herpera.com/mhdfrog.htm （页面存档备份，存于互联网档案馆）

[1] Frost, Darrel R. 2023. Amphibian Species of the World: an online reference. Version 6.1 (21 March 2023). Electronic Database accessible at https://amphibiansoftheworld.amnh.org/Amphibia/Anura/Megophryidae/Megophryinae/Megophrys （页面存档备份，存于互联网档案馆）. American Museum of Natural History, New York, USA.

[Mahony-2] 2.0 ^2.1 ^2.2 ^2.3 ^2.4 ^2.5 ^2.6 Mahony, Stephen, Emma C. Teeling & S. D. Biju. 2013. Three New Species of Horned Frogs, Megophrys (Amphibia: Megophryidae), from Northeast India, with a resolution to the identity of Megophrys boettgeri populations reported from the region. Zootaxa. 3722(2): 143-169.

[3] Truong Quang Nguyen, Cuong The Pham, Tao Thien Nguyen, Anh Mai Luong and Thomas Ziegler. 2020. A New Species of Megophrys (Amphibia: Anura: Megophryidae) from Vietnam. Zootaxa. 4722(5); 401–422. DOI: 10.11646/zootaxa.4722.5.1

[Wang-4] 4.0 ^4.1 Yingyong Wang, Jian Zhao, Jianhuan Yang, Zhixin Zhou, Guoling Chen and Yang Liu. 2014. Morphology, Molecular Genetics, and Bioacoustics Support Two New Sympatric Xenophrys Toads (Amphibia: Anura: Megophryidae) in Southeast China. PLoS ONE. 9(4): e93075. DOI: 10.1371/journal.pone.0093075

[5] Benjamin Tapley, Timothy Cutajar, Stephen Mahony, Chung T. Nguyen, Vinh Q. Dau, Anh M. Luong, Dzung T. Le, Tao T. Nguyen, Truong Q. Nguyen, Christopher Portway, Hào V. Lương and Jodi J. L. Rowley. 2018. Two New and Potentially Highly threatened Megophrys Horned Frogs (Amphibia: Megophryidae) from Indochina’s Highest Mountains. Zootaxa. 4508(3); 301–333. DOI: 10.11646/zootaxa.4508.3.1

[6] Benjamin Tapley, Timothy Cutajar, Luan Thanh Nguyen, Christopher Portway, Stephen Mahony, Chung Thanh Nguyen, Luke Harding, Hao Van Luong and Jodi J. L. Rowley. 2020. A New Potentially Endangered Species of Megophrys (Amphibia: Megophryidae) from Mount Ky Quan San, north-west Vietnam. Journal of Natural History. 54(39-40); 2543-2575. DOI: 10.1080/00222933.2020.1856952

[7] Stephen Mahony, Rachunliu G. Kamei, Emma C. Teeling and S. D. Biju. 2018. Cryptic Diversity within the Megophrys major Species Group (Amphibia: Megophryidae) of the Asian Horned Frogs: Phylogenetic Perspectives and A Taxonomic Revision of South Asian Taxa, with Descriptions of Four New Species. Zootaxa. 4523(1); 1–96. DOI: 10.11646/zootaxa.4523.1.1

[8] Wang, Ying-yong, Tian-du Zhang, Jian Zhao, Yik-Hei Sung, Jian-huan Yang, Hong Pang & Zhong Zhang. 2012. Description of A New Species of the Genus Xenophrys Günther, 1864 (Amphibia: Anura: Megophryidae) from Mount Jinggang, China, based on Molecular and Morphological data. Zootaxa. (3546): 53-67.

[9] Misbahul Munir, Amir Hamidy, Masafumi Matsui, Djoko T. Iskandar, Irvan Sidik and Tomohiko Shimada. 2019. A New Species of Megophrys Kuhl & Van Hasselt (Amphibia: Megophryidae) from Borneo allied to M. nasuta (Schlegel, 1858). Zootaxa. 4679(1); 1–24. DOI: 10.11646/zootaxa.4679.1.1

[10] Misbahul Munir, Amir Hamidy, Achmad Farajallah and Eric N. Smith. 2018. A New Megophrys Kuhl and Van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa. 4442(3); 389–412. DOI: 10.11646/zootaxa.4442.3.3

[11] 云南高黎贡山发现两栖动物一新种――泸水角蟾. 发现. [2023-07-12]. （原始内容存档于2023-07-12）.

[12] Stephen Mahony, Rachunliu G. Kamei, Emma C. Teeling and S. D. Biju. 2020. Taxonomic Review of the Asian Horned Frogs (Amphibia: Megophrys Kuhl & Van Hasselt) of Northeast India and Bangladesh previously misidentified as M. parva (Boulenger), with Descriptions of Three New Species. Journal of Natural History. 54(1-4 - Amphibian taxonomy: Early 21st century case studies); DOI: 10.1080/00222933.2020.1736679

[13] Kevin R. Messenger, Hollis A. Dahn, Yuanrui Liang, Peng Xie, Yong Wang and Changhu Lu. 2019. A New Species of the Genus Megophrys Gunther, 1864 (Amphibia: Anura: Megophryidae) from Mount Wuyi, China. Zootaxa. 4554(2); 561–583. DOI: 10.11646/zootaxa.4554.2.9

[14] Benjamin Tapley, Timothy Cutajar, Stephen Mahony, Chung Thanh Nguyen, Vinh Quang Dau, Tao Thien Nguyen, Hao Van Luong and Jodi J. L. Rowley. 2017. The Vietnamese Population of Megophrys kuatunensis (Amphibia: Megophryidae) Represents A New Species of Asian Horned Frog from Vietnam and southern China. Zootaxa. 4344(3); 465–492. DOI: 10.11646/zootaxa.4344.3.3

[15] Misbahul Munir, Kanto Nishikawa, Amir Hamidy and Eric N. Smith. 2021. Two New Species of Megophrys Kuhl and Van Hasselt (Amphibia: Megophryidae) from Sumatra, Indonesia. Zootaxa. 5057(4); 503-529. DOI: 10.11646/zootaxa.5057.4.3

[16] Kai, Wang; Jinlong, Ren; Hongman, Chen; Zhitong, Lyu; Xianguang, Guo; Ke, Jiang; Jinmin, Chen; Jiatang, Li; Peng, Guo; Yingyong, Wang; Jing, Che. The updated checklists of amphibians and reptiles of China. Biodiversity Science. 2020-02-20, 28 (2): 189. doi:10.17520/biods.2019238.

[17] Chen, Jin-Min; Poyarkov, Nikolay A.; Suwannapoom, Chatmongkon; Lathrop, Amy; Wu, Yun-He; Zhou, Wei-Wei; Yuan, Zhi-Yong; Jin, Jie-Qiong; Chen, Hong-Man; Liu, He-Qun; Nguyen, Truong Quang; Nguyen, Sang Ngoc; Duong, Tang Van; Eto, Koshiro; Nishikawa, Kanto; Matsui, Masafumi; Orlov, Nikolai L.; Stuart, Bryan L.; Brown, Rafe M.; Rowley, Jodi J.L.; Murphy, Robert W.; Wang, Ying-Yong; Che, Jing. Large-scale phylogenetic analyses provide insights into unrecognized diversity and historical biogeography of Asian leaf-litter frogs, genus Leptolalax (Anura: Megophryidae). Molecular Phylogenetics and Evolution. 2018-07, 124: 162–171. doi:10.1016/j.ympev.2018.02.020.

[18] Mahony, Stephen; Foley, Nicole M.; Biju, S.D.; Teeling, Emma C. Evolutionary History of the Asian Horned Frogs (Megophryinae): Integrative Approaches to Timetree Dating in the Absence of a Fossil Record. Molecular Biology and Evolution. 2017-01-18: msw267. doi:10.1093/molbev/msw267.

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