Trochoidea (superfamily)

Trochoidea; Temporal range: Ordovician–Recent PreꞒ Ꞓ O S D C P T J K Pg N
	A live individual of Tegula pulligo
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Vetigastropoda
Order:	Trochida
Superfamily:	Trochoidea; Rafinesque, 1815
Families
	See text
Synonyms
	Angarioidea Gray, 1857; Phasianelloidea Swainson, 1840; Trochacea (suffix -oidea mandatory for a superfamily name following current ICZN art. 29.2.); Turbinoidea Rafinesque, 1815;

Trochoidea is a superfamily of small to very large vetigastropod sea snails with gills and an operculum.^[2] Species within this superfamily have nacre as the inner shell layer. The families within this superfamily include the Trochidae, the top snails. This superfamily is the largest vetigastropodan superfamily, containing more than 2,000 species.^[3]

This taxon is not the same as a pulmonate land snail genus which is spelled the same way: Trochoidea (genus).

Taxonomy

2005 taxonomy

This superfamily consisted of nine following families (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):

Trochidae Rafinesque, 1815
Calliostomatidae Thiele, 1924 (1847)
† Elasmonematidae Knight, 1956
† Eucochlidae Bandel, 2002
† Microdomatidae Wenz, 1938
† Proconulidae Cox, 1960
Solariellidae Powell, 1951 - synonym: Minoliinae Kuroda, Habe & Oyama, 1971
† Tychobraheidae Horný, 1992
† Velainellidae Vasseur, 1880

2017 taxonomy

Williams et al. (2008)^[4] have made further refinements to the taxonomy of Trochoidea. The revised taxonomy as defined by Bouchet et al. (2017) resulted in the following families.^[5] The fossil families, tentatively included in this list, are based on Tracey et al. (1993) ^[6]

Angariidae Gray, 1857
† Anomphalidae Wenz, 1938
† Araeonematidae Nützel, 2012
Areneidae McLean, 2012
Calliostomatidae Thiele, 1924 (1847)
Colloniidae Cossmann, 1917
Conradiidae Golikov & Starobogatov, 1987
† Elasmonematidae Knight, 1956
† Epulotrochidae Gründel, Keupp & Lang, 2017
† Eucochlidae Bandel, 2002
Liotiidae Gray, 1850
Margaritidae Thiele, 1924
†Metriomphalidae Gründel, Keupp & Lang, 2017
† Microdomatidae Wenz, 1938
†Nododelphinulidae Cox, 1960
Phasianellidae Swainson, 1840
† Proconulidae Cox, 1960
† Sclarotrardidae Gründel, Keupp & Lang, 2017
Skeneidae Clark W., 1851
Solariellidae Powell, 1951
Tegulidae Kuroda, Habe & Oyama, 1971
Trochidae Rafinesque, 1815
Turbinidae Rafinesque, 1815
† Tychobraheidae Horný, 1992
† Velainellidae Vasseur, 1880
Trochoidea (unassigned)

Unassigned to a family:

Margarella Thiele, 1893 - Margarella rosea belongs to Cantharidinae,^[7] but Margarella antarctica belongs to Calliostomatidae/Thysanodontinae as Carinastele antarctica.^[7]

Nomen dubium

Ataphridae Cossmann, 1915

Families brought into synonymy

Crosseolidae Hickman, 2013: synonym of Conradiidae Golikov & Starobogatov, 1987
Cyclostrematidae P. Fischer, 1865: synonym of Liotiidae Gray, 1850
Delphinulidae Stoliczka, 1868: synonym of Angariidae Gray, 1857
Gazidae Hickman & McLean, 1990: synonym of Margaritidae Thiele, 1924
Minoliinae Kuroda, Habe & Oyama, 1971: synonym of Solariellidae Powell, 1951
† Parataphrinae Calzada, 1989: synonym of † Proconulidae Cox, 1960
Stomatellidae Gray, 1840: synonym of Stomatellinae Gray, 1840
Stomatiidae Carpenter, 1861: synonym of Stomatellinae Gray, 1840
Tricoliidae Woodring, 1928: synonym of Tricoliinae Woodring, 1928 (rank change)

References

^ Lindberg, edited by Winston F. Ponder, David R. (2008). Phylogeny and evolution of the Mollusca. Berkeley: University of California Press. ISBN 978-0-520-25092-5. {{cite book}}: |first1= has generic name (help)CS1 maint: multiple names: authors list (link)
^ Gofas, S. (2013). Trochoidea. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=156489 on 2013-06-29
^ Williams, Suzanne T. (2012). "Advances in molecular systematics of the vetigastropod superfamily Trochoidea". Zoologica Scripta. 41 (6): 571–595. doi:10.1111/j.1463-6409.2012.00552.x.
^ Williams S. T., Karube S. & Ozawa T. (September 2008) "Molecular systematics of Vetigastropoda: Trochidae, Turbinidae and Trochoidea redefined". Zoologica Scripta 37(5): 483-506. doi:10.1111/j.1463-6409.2008.00341.x
^ "Revised Classification, Nomenclator and Typification of Gastropod and Monoplacophoran Families" by Philippe Bouchet & Jean-Pierre Rocroi, Bernhard Hausdorf, Andrzej Kaim, Yasunori Kano, Alexander Nützel, Pavel Parkhaev, Michael Schrödl and Ellen E. Strong in Malacologia, 2017, 61(1–2): 1–526
^ Tracey S., J.A. Todd & D.H. Erwin, 1993, Mollusca: Gastropoda. Pp. 131–167, in: M. J. Benton, ed., The Fossil Record, volume 2. chapman & hall, london. 845 pp.
^ ^a ^b Williams S.T., Donald K.M., Spencer H.G. & Nakano T. (2010) Molecular systematics of the marine gastropod families Trochidae and Calliostomatidae (Mollusca: Superfamily Trochoidea). Molecular Phylogenetics and Evolution 54:783-809.