Psocomorpha

Psocomorpha Temporal range: Cenomanian–Recent PreꞒ Ꞓ O S D C P T J K Pg N
Abelopsocus basipunctatus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
(unranked):	Paraneoptera
Order:	Psocodea
Suborder:	Psocomorpha Roesler, 1944
Infraorders[1]
Archipsocetae Caeciliusetae Epipsocetae Homilopsocidea Philotarsetae Psocetae

Psocomorpha is a suborder of barklice, booklice, and parasitic lice in the order Psocodea (formerly Psocoptera).[2][3][1] There are more than 20 families and 5,300 described species in Psocomorpha.[4][5][6]

Xanthocaecilius sommermanae

Phylogeny

The below cladogram of Psocodea shows the position of Psocomorpha:[1]

Psocodea	Troctomorpha Phthiraptera Philopteridae Anoplura Rhynchophthirina Trichodectidae Amblycera Liposcelididae Pachytroctidae Sphaeropsocidae Amphientometae Psocomorpha Homilopsocidea Caeciliusetae Psocetae Epipsocetae Philotarsetae Archipsocetae Trogiomorpha Atropetae Psyllipsocetae Prionoglaridetae (paraphyletic)

Fossil record

The oldest fossil record of the suborder is Burmesopsocus lienhardi from the mid-Cretaceous (early Cenomanian) Burmese amber, which is not assigned to any family.[7]

Classification

Psocomorpha contains 6 infraorders and 26 families:[8]

• Archipsocetae
  • Archipsocidae Pearman, 1936 (ancient barklice)
• Caeciliusetae
  • Amphipsocidae Pearman, 1936 (hairy-winged barklice)
  • Asiopsocidae Mockford & Garcia Aldrete, 1976
  • Caeciliusidae Mockford, 2000 (lizard barklice)
  • Dasydemellidae Mockford, 1978 (shaggy psocids)
  • Paracaeciliidae Mockford, 1989
  • Stenopsocidae Pearman, 1936 (narrow barklice)
• Epipsocetae
  • Cladiopsocidae Smithers, 1972
  • Dolabellopsocidae Eertmoed, 1973
  • Epipsocidae Pearman, 1936 (elliptical barklice)
  • Ptiloneuridae Roesler, 1940
  • Spurostigmatidae Eertmoed, 1973
• Homilopsocidea
  • Ectopsocidae Roesler, 1944 (outer barklice)
  • Elipsocidae Pearman, 1936 (damp barklice)
  • Lachesillidae Pearman, 1936 (fateful barklice)
  • Lesneiidae Schmidt & New, 2004
  • Mesopsocidae Pearman, 1936 (middle barklice)
  • Peripsocidae Roesler, 1944 (stout barklice)
  • Sabulopsocidae Schmidt & New, 2004
• Philotarsetae
  • Philotarsidae Pearman, 1936 (loving barklice)
  • Pseudocaeciliidae Pearman, 1936 (false lizard barklice)
  • Trichopsocidae Pearman, 1936 (lash-faced psocids)
• Psocetae
  • Hemipsocidae Pearman, 1936 (leaf litter barklice)
  • Myopsocidae Pearman, 1936 (mouse-like barklice)
  • Psilopsocidae Roesler, 1944
  • Psocidae Hagen, 1865 (common barklice)

References

1. de Moya, Robert S; Yoshizawa, Kazunori; Walden, Kimberly K O; Sweet, Andrew D; Dietrich, Christopher H; Kevin P, Johnson (2021-06-16). Buckley, Thomas (ed.). "Phylogenomics of Parasitic and Nonparasitic Lice (Insecta: Psocodea): Combining Sequence Data and Exploring Compositional Bias Solutions in Next Generation Data Sets". Systematic Biology. 70 (4): 719–738. doi:10.1093/sysbio/syaa075. ISSN 1063-5157. PMID 32979270.
2. Johnson, Kevin P.; Smith, Vincent S. (2019). "Psocodea Species File Online". Retrieved 2019-06-14.
3. Bess, Emilie; Smith, Vince; Lienhard, Charles; Johnson, Kevin P. (2006). "Psocodea". Tree of Life. Retrieved 2019-06-14.
4. "Psocomorpha Report". Integrated Taxonomic Information System. Retrieved 2019-05-04.
5. "Psocomorpha suborder Information". BugGuide.net. Retrieved 2019-05-04.
6. Johnson, Kevin P.; Smith, Vincent S. (2019). "suborder Psocomorpha". Psocodea species file online, Version 5.0. Retrieved 2019-05-04.
7. Yoshizawa, Kazunori; Yamamoto, Shuhei (October 2021). "The earliest fossil record of the suborder Psocomorpha (Insecta: Psocodea) from mid-Cretaceous Burmese amber, with description of a new genus and species". Insecta Matsumurana. New Series. 77: 1–15. ISSN 0020-1804.
8. Johnson, Kevin P.; Smith, Vincent S. (2021). "Psocodea species file online, Version 5.0". Retrieved 2021-11-05.

Further reading

• Lyal, Ch H. C. (1985). "Phylogeny and classification of the Psocodea, with particular reference to the lice (Psocodea: Phthiraptera)". Systematic Entomology. 10 (2): 145–165. Bibcode:1985SysEn..10..145L. doi:10.1111/j.1365-3113.1985.tb00525.x. ISSN 0307-6970. S2CID 86331606.
• Yoshizawa, Kazunori (2002). "Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea: 'Psocoptera')". Zoological Journal of the Linnean Society. 136 (3): 371–400. doi:10.1046/j.1096-3642.2002.00036.x. ISSN 0024-4082.
• Yoshizawa, Kazunori (2005). "Morphology of Psocomorpha (Psocodea:'Psocoptera')". Insecta Matsumurana. New Series. 62: 1–44. hdl:2115/10524.

External links

• Media related to Psocomorpha at Wikimedia Commons