Characteristics include partial to complete division of the eyes by a canthus, geniculateantennae, and distinctly separated coxae. The body is typically elongated and slightly flattened.
G. Cassis, W.W.K. Houston, T.A. Weir & B.P. Moore (1992). Updated by A.A. Calder (2002). Coleoptera: Scarabaeoida. Australian Faunal Directory. Australian Biological Resources Study, Canberra. Viewed 22 November 2007.