Leptopleuroninae

Leptopleuroninae Temporal range: Early — Late Triassic
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Clade:	†Parareptilia
Order:	†Procolophonomorpha
Family:	†Procolophonidae
Subfamily:	†Leptopleuroninae Ivakhnenko, 1979
Genera
†Basileosaurus? †Cornualbus[1] †Hwiccewyrm †Hypsognathus †Koiloskiosaurus †Leptopleuron †Libognathus †Mandaphon †Neoprocolophon †Pentaedrusaurus †Phonodus †Sclerosaurus †Scoloparia †Soturnia †Threordatoth

Leptopleuroninae is an extinct subfamily of procolophonid reptiles.[2] It is defined as all taxa closer to Leptopleuron lacertinum than to Procolophon trigoniceps.[2] The oldest member of Leptopleuroninae is Phonodus dutoitorum from the Induan age of the Early Triassic.[3] It is the only procolophonid group that survived into the Late Triassic.[4]

Phylogeny

A cladogram showing relationships within Procolophonidae after Modesto et al., 2010:[3]

Procolophonidae	Coletta seca Pintosaurus magnidentis Sauropareion anoplus Kitchingnathus untabeni Phaanthosaurus sp. Theledectinae Tichvinskia vjatkensis Procolophoninae Leptopleuroninae Pentaedrusaurus ordosianus Neoprocolophon asiaticus Phonodus dutoitorum clade "N" Sclerosaurus armatus Scoloparia glyphanodon Leptopleuron lacertinum Soturnia caliodon Hypsognathus fenneri

Below are two cladograms that follow phylogenetic analyses by Butler et al. (2023):[4]

Analyses 1 and 3: Strict consensus of 760 and 18 most parsimonious trees (MPTs). Leptopleuroninae Neoprocolophon Pentaedrusaurus Mandaphon Scoloparia Sclerosaurus Hwiccewyrm Hypsognathus Soturnia Leptopleuron

Analysis 2: Single MPT. Leptopleuroninae Kapes bentoni Thelephon Kapes majmesculae Pentaedrusaurus Anomoiodon Theledectes Procolina Phonodus Mandaphon Neoprocolophon Scoloparia Sclerosaurus Hwiccewyrm Leptopleuron Hypsognathus Soturnia

References

1. Silva-Neves, E.; Da-Rosa, Á. A. S.; Modesto, S. P.; Dias-da-Silva, S. (2024). "Cornualbus primus gen. et sp. nov.: a new procolophonid (Reptilia: Parareptilia) from Upper Triassic of South America, first tetrapod from the Passo das Tropas Member of the Santa Maria Supersequence". Journal of Systematic Palaeontology. 22 (1). doi:10.1080/14772019.2024.2373116.
2. Cisneros, J. C. (2008). "Phylogenetic relationships of procolophonid parareptiles with remarks on their geological record". Journal of Systematic Palaeontology. 6 (3): 345–366. doi:10.1017/S1477201907002350. S2CID 84468714.
3. Modesto, S.P.; Scott, D.M.; Botha-Brink, J.; Reisz, R.R. (2010). "A new and unusual procolophonid parareptile from the Lower Triassic Katberg Formation of South Africa". Journal of Vertebrate Paleontology. 30 (3): 715–723. Bibcode:2010JVPal..30..715M. doi:10.1080/02724631003758003. S2CID 84563475.
4. Butler, R. J.; Meade, L. E.; Cleary, T. J.; McWhirter, K. T.; Brown, E. E.; Kemp, T. S.; Benito, J.; Fraser, N. C. (2023). "Hwiccewyrm trispiculum gen. et sp. nov., a new leptopleuronine procolophonid from the Late Triassic of southwest England". The Anatomical Record. doi:10.1002/ar.25316. PMID 37735997.