It was originally placed within the Graphidaceae family and subfamily Redonographoideae.Lücking, Tehler & Lumbsch (2013).[5][6][7]
Molecular clock analysis indicates that Redonographoideae diverged from Graphidoideae subfamily about 132 million years ago (Ma) in the Early Cretaceous age.[5]
However, Kraichak et al. in 2018,[8] ranked Graphidales as a separate order based on a temporal approach, and accepted five separate families; Diploschistaceae, Fissurinaceae, Gomphillaceae, Graphidaceae and Thelotremataceae.[9][10] Wijayawardene et al. 2020 agreed but also added family Redonographaceae to the order, which included Gymnographopsis.[11][2]
^ abLücking, R.; Tehler, A.; Bungartz, F.; Rivas Plata, E.; Lumbsch, H.T. (2013). "Journey from the West: Did tropical Graphidaceae (lichenized Ascomycota: Ostropales) evolve from a saxicolous ancestor along the American Pacific coast?". American Journal of Botany. 100 (5): 844–856. doi:10.3732/ajb.1200548. PMID23594913.
^Lücking, Robert; Hodkinson, Brendan P.; Leavitt, Steven D. (2017). "The 2016 classification of lichenized fungi in the Ascomycota and Basidiomycota–Approaching one thousand genera". The Bryologist. 119 (4): 361–416. doi:10.1639/0007-2745-119.4.361. S2CID90258634.
^Miranda-González, Ricardo; Lücking, Robert; Barcenas-Peña, Alejandrina; Herrera-Campos, María de los Ángeles (2020). "The new genus Jocatoa (Lecanoromycetes: Graphidaceae) and new insights into subfamily Redonographoideae". The Bryologist. 123 (2): 127–143. doi:10.1639/0007-2745-123.2.127. S2CID215812045.
^Kraichak, Ekaphan; Huang, Jen-Pan; Nelsen, Matthew; Leavitt, Steven D.; Lumbsch, H. Thorsten (2018). "A revised classification of orders and families in the two major subclasses of Lecanoromycetes (Ascomycota) based on a temporal approach". Botanical Journal of the Linnean Society. 188: 233–249. doi:10.1093/botlinnean/boy060.
^Behera, Prashant Kumar; Nayaka, Sanjeeva (April 2021). "New Distributional Records to Lichen Biota of Assam, India". Indian Forester. 147 (4): 400–404. doi:10.36808/if/2021/v147i4/152523. S2CID247340504.
^Follmann, G. (1967). "Die Flechtenflora der nordchilenischen Nebeloase Cerro Moreno". Nova Hedwigia (in German). 14: 213–281.
^Miranda-González, Ricardo; Lücking, Robert; Barcenas-Peña, Alejandrina; Ángeles Herrera-Campos, María de los (2020). "The new genus Jocatoa (Lecanoromycetes: Graphidaceae) and new insights into subfamily Redonographoideae". The Bryologist. 123 (2): 127–143. doi:10.1639/0007-2745-123.2.127. S2CID215812045.
^Egea, J.M.; Torrente, P. (1996). "Tres nuevas especies de hongos liquenizados de la Provincia del Cabo (Sudáfrica)". Cryptogamie Bryologie Lichénologie (in Spanish). 17 (4): 295–312.