^Matos-Maraví P.F., Peña C., Willmott K.R., Freitas A.V.L., Wahlberg N. 2012. Systematics and evolutionary history of butterflies in the “Taygetis clade” (Nymphalidae: Satyrinae: Euptychiina): Towards a better understanding of Neotropical biogeography. Molecular Phylogenetics and Evolution. 66:54–68. doi:10.1016/j.ympev.2012.09.005
^Freitas, A.V.L. & C. Peña. 2006. Description of genus Guaianaza for "Euptychia" pronophila (Lepidoptera: Nymphalidae: Satyrinae) with a description of the immature stages. Zootaxa 1163: 49-59.[1]