Galacticidae is a recently recognised and enigmatic family of insects in the lepidopteranorder. These moderate sized moths are 8–17 mm in wingspan and have previously been embedded within several lepidopteran superfamilies (Tineoidea: Psychidae, Urodoidea, Sesioidea and in several families of Yponomeutoidea), but Galacticidae is currently placed in its own superfamily at the base of the natural group Apoditrysia (Dugdale et al., 1999 [1998]; May, 2004).
Note: the superfamily was unintentionally called "Galaticoidea" in Dugdale et al. (1999) [1998].
The relationships of Galacticidae need reassessment with new characters including DNA data [1]. The genus Homadaula looks remarkably like the yponomeutid genera Prays and Atemelia and some species are reminiscent of "small ermine" moths. Despite the spined abdominal segments of the pupae and a few other characters (Minet, 1986) some possess structural similarities to yponomeutids as well and have similar larval behaviour [2] so their removal from the Yponomeutoidea has been questioned (Mey, 2004).
Dugdale, J.S., Kristensen, N.P., Robinson, G.S. and Scoble, M.J. (1999) [1998]. The smaller microlepidoptera grade superfamilies, Ch.13., pp. 217–232 in Kristensen, N.P. (Ed.). Lepidoptera, Moths and Butterflies. Volume 1: Evolution, Systematics, and Biogeography. Handbuch der Zoologie. Eine Naturgeschichte der Stämme des Tierreiches / Handbook of Zoology. A Natural History of the phyla of the Animal Kingdom. Band / Volume IV Arthropoda: Insecta Teilband / Part 35: 491 pp. Walter de Gruyter, Berlin, New York.
Mey, W. (2004). Galacticidae (Ditrysia incertae sedis). Esperiana1: 91-102.
Minet, J. (1986). Ébauche d'une classification moderne de l'ordre des Lépidoptères. Alexanor 14(7): 291-313.
Moriuti, S. (1963). Remarks on the Paraprays anisocentra (Meyrick, 1922) (Plutellidae), with descriptions of its larva and pupa. Transactions of the Lepidopterological Society of Japan, 14(3): 52-59.