Kovalevsky discovered that tunicates are not molluscs, but that their larval stage has a notochord and pharyngeal slits, like vertebrates. Further, these structures develop from the same germ layers in the embryo as the equivalent structures in vertebrates, so he argued that the tunicates should be grouped with the vertebrates as chordates. 19th-century zoology thus converted embryology into an evolutionary science, connecting phylogeny with homologies between the germ layers of embryos, foreshadowing evolutionary developmental biology.[5]